Éditeur(s) : J.W. Markham & A.L. Duda, pp. 31-59. Fort Pierce, FL : IAMSLIC
Résumé : A time when France, as well as Great Britain, had a very ambitious maritime policy, the La Pérouse expedition was the only maritime expedition conceived, designed and followed at a national level by the King himself, Louis XVI. The excellence of the Expedition's preparation and of its importance in terms of scientific results, the exceptional concern of La Pérouse in the fields of data gathering and transmission, his very high quality as captain of a global expedition, made it a remarkable achievement, despite its disastrous end.

Droits : info:eu-repo/semantics/openAccess
http://archimer.ifremer.fr/doc/2004/acte-358.pdf
http://archimer.ifremer.fr/doc/00000/358/

Éditeur(s) : HAL CCSD Public Library of Science
Résumé : International audience
The giant bathymodioline mussels from vents have been studied as models to understand the adaptation of organisms to deep-sea chemosynthetic environments. These mussels are closely related to minute mussels associated to organic remains decaying on the deep-sea floor. Whereas biological data accumulate for the giant mussels, the small mussels remain poorly studied. Despite this lack of data for species living on organic remains it has been hypothesized that during evolution, contrary to their relatives from vents or seeps, they did not acquire highly specialized biological features. We aim at testing this hypothesis by providing new biological data for species associated with organic falls. Within Bathymodiolinae a close phylogenetic relationship was revealed between the Bathymodiolus sensu stricto lineage (i.e. ``thermophilus'' lineage) which includes exclusively vent and seep species, and a diversified lineage of small mussels, attributed to the genus Idas, that includes mostly species from organic falls. We selected Idas iwaotakii (Habe, 1958) from this latter lineage to analyse population structure and to document biological features. Mitochondrial and nuclear markers reveal a north-south genetic structure at an oceanic scale in the Western Pacific but no structure was revealed at a regional scale or as correlated with the kind of substrate or depth. The morphology of larval shells suggests substantial dispersal abilities. Nutritional features were assessed by examining bacterial diversity coupled by a microscopic analysis of the digestive tract. Molecular data demonstrated the presence of sulphur-oxidizing bacteria resembling those identified in other Bathymodiolinae. In contrast with most Bathymodiolus s.s. species the digestive tract of I. iwaotakii is not reduced. Combining data from literature with the present data shows that most of the important biological features are shared between Bathymodiolus s.s. species and its sister-lineage. However Bathymodiolus s.s. species are ecologically more restricted and also display a lower species richness than Idas species.
ISSN: 1932-6203

hal-01545914
https://hal.archives-ouvertes.fr/hal-01545914
DOI : 10.1371/journal.pone.0069680

Éditeur(s) : HAL CCSD Elsevier
Résumé : International audience
The present study is based on a set of lavas and crosscutting dikes collected by dives along detailed vertical transects on the northern flank of the western part of the Blanco Transform Fault, Northeast Pacific. The studied area consists of a small basin, the Western Blanco Depression (WBD), extending from the southern end of the Juan de Fuca ridge to a pseudofault trace 60 km eastward. The Northern Scarp of the WBD comprises a volcanic unit overlying a sheeted-dike complex. Major and trace element data, coupled with Sr–Nd isotope ratios, reveal a two-component mantle source, composed by an isotopically depleted matrix variably veined by more enriched material. One chemical group (NS2), indistinguishable from the other Northern Scarp samples on the basis of trace element data, has an unusually depleted isotopic composition typical of a nearly pure mantle end-member. Some cogenetic samples of the Northern Scarp have been used to constrain the differentiation modalities. Anorthite and MgO content profiles in plagioclase xenocrysts and phenocrysts reveal (i) the existence of H2O-bearing evolved melts in the mushy zones and (ii) the occurrence of mixing process between these melts and anhydrous mafic liquids. The hydration is supported by other petrographic features such as high magmatic fO2 values, calculated from Fe–Ti oxide pairs, and the presence of pyroxene inclusions in plagioclase phenocrysts. Mixing, consistent with the existence of Ni-rich ferrobasalts, is interpreted to be the consequence of the reservoir refilling by mafic liquids (Mg# = 70). These petrological and geochemical evidences are combined with the evolution of Mg# with depth to suggest a periodic open-system magma chamber evolution beneath the southern end of the Juan de Fuca ridge.
ISSN: 0024-4937

insu-00213426
https://hal-insu.archives-ouvertes.fr/insu-00213426
DOI : 10.1016/j.lithos.2007.06.009

Éditeur(s) : HAL CCSD Elsevier
Résumé : International audience
A comprehensive inventory of global biodiversity would be greatly improved by automating methods for species delimitation. The Automatic Barcode Gap Discovery method, the Poisson tree processes algorithm and the Generalized mixed Yule coalescent model have been proposed as means of increasing the rate of biodiversity description using single locus data. We applied these methods to explore the diversity within the Aglaopheniidae, a hydrozoan family with many species widely distributed across tropical and tem perate oceans. Our analyses revealed widespread cryptic diversity in this family, almost half of the morpho species presenting several independent evolutionary lineages, as well as support for cases of synonymy. For two common species of this family, Lytocarpia brevirostris and Macrorhynchia phoenicea, we compared the outputs to clustering analyses based on microsatellite data and to nuclear gene phylo genies. For L. brevirostris, microsatellite data were congruent with results of the species delimitation methods, revealing the existence of two cryptic species with Indo Pacific distribution. For M. phoenicea, all analyses confirmed the presence of two cryptic species within the South Western Indian Ocean. Our study suggests that the diversity of Aglaopheniidae might be much higher than assumed, likely related to low dispersal capacities. Sequence based species delimitation methods seem highly valuable to reveal cryptic diversity in hydrozoans; their application in an integrative framework will be very useful in describing the phyletic diversity of these organisms.
ISSN: 1055-7903

hal-01359407
https://hal.archives-ouvertes.fr/hal-01359407
https://hal.archives-ouvertes.fr/hal-01359407/document
https://hal.archives-ouvertes.fr/hal-01359407/file/Postaire-etal2016MolPhylEvol_sp_delim_hydraires_HAL.pdf
DOI : 10.1016/j.ympev.2016.08.013

Éditeur(s) : HAL CCSD Elsevier
Résumé : International audience
Periodically harvested closures (PHCs) are small fisheries closures with objectives such as sustaining fisheriesand conserving biodiversity and have become one of the most common forms of nearshore marine managementin the Western Pacific. Although PHCs can provide both short-termconservation and fisheries benefits, their potentialas a long-term management strategy remains unclear. Through empirical assessment of a single harvestevent in each of five PHCs, we determined whether targeted fishes that differ in their vulnerability to fishing recoveredto pre-harvest conditions (the state prior to last harvest) and demonstrated post-harvest recovery benefitsafter 1 year of re-closure. For low and moderately vulnerable species, two PHCs provided significant preharvestbenefits and one provided significant post-harvest recovery benefits, suggesting a contribution to longer-term sustainability. PHCs with a combination of high compliance and longer closing times are more likelyto provide fisheries benefits and recover from harvest events, however, no benefits were observed across anyPHCs for highly vulnerable species. We recommend PHCs have longer closure periods before being harvestedand species that are highly vulnerable to fishing (e.g. large species of; grouper,wrasse and parrotfish) are avoidedduring harvests to avoid overexploitation and increase the sustainability of small-scale fisheries.
ISSN: 0006-3207

hal-01381316
https://hal-univ-perp.archives-ouvertes.fr/hal-01381316
DOI : 10.1016/j.biocon.2016.08.038

Éditeur(s) : Gauthier-Villars
Résumé : Since the advent of plate tectonics the active margins of the oceans have been studied according to a model whose basis is a piling up of imbrications of oceanic material units, in front of the upper plate, forming a great tectonic accretionary prism. This model is broadly the same for all the authors; it has been largely used for alpine-type or paleo-oceanic mountain chain, all around the world. The authors have directly taken part in the drillings on western and eastern Pacific active margins (Legs 60 and 67) and they have worked on the tectonics of alpine (paleo-oceanic) chains of the peri-mediterranean, peri-caribbean, and peripacific belts. They compare the geological data acumulated from these tow direct sources. The tectonic constraints constraints regime is specified and the consequences of the subduction are looked for. The notions of collison and obduction are discussed in relation to subduction and tectonics. Finally, an anlaysis of the temporal evolution of active margins and of chains, shows the necessity of thinking in terms of superimposed tectonics.
Oceanologica Acta, Special issue (0399-1784) (Gauthier-Villars), 1981

Droits : info:eu-repo/semantics/openAccess
http://archimer.ifremer.fr/doc/00246/35700/34207.pdf
http://archimer.ifremer.fr/doc/00246/35700/

Éditeur(s) : HAL CCSD Elsevier
Résumé : International audience
Geochronological studies of samples from continuous crustal sections of fossil intra-oceanic arcs are paramount for determining the precise timing of major processes that took place during the building of an oceanic island-arc crust. In this study, laser ablation ICP-MS U-Pb zircon (and conventional Rb-Sr) analyses pinpoint the timing of major events responsible for crustal growth of the Kohistan paleo-island arc (Northern Pakistan). Inheritance in magmatic rocks is evidence for recycling of intra-arc material and indicates that the Kohistan arc may have formed as early as 135 +/- 4 Ma. One older inherited grain indicates that the arc developed on a young oceanic lithosphere whose remnants include a c. 175 Ma old component. An interesting consequence is that intra-oceanic arc magmas can yield inherited zircons, which can be much older than the arc system itself without requiring recycling back into the mantle by subduction processes. In the case of the Kohistan arc, ante-arc Neotethys oceanic lithosphere was tapped by arc magmas during their way upward into the arc crust. The oldest magmatic ages measured in this study fall in the range 101-102 Ma (100.9 +/- 0.6 and 102.1 +/- 0.4 Ma). This period corresponds to arc build-up and thickening of the arc crust. Leucogranitic melts dated at 89.9 +/- 0.4 Ma and 90.9 +/- 1.0 Ma are considered as produced by dehydration/melting reaction accompanying granulitisation of the thickened lower arc crust. A maximum age for this event is 97.7 +/- 0.7 Ma, the age of emplacement of a garnet meta-tonalite affected by granulite facies metamorphism. Magmatic activity was still ongoing during and after the granulitisation as testified by emplacement of a diorite and a gabbro dated at 89.1 +/- 0.6 and 88.2 +/- 0.9 Ma respectively. The upper part of the metaplutonic sequence contains diorite samples dated at 84.6 +/- 0.5 Ma and 84.3 +/- 0.5 Ma and as young as 81.1 +/- 0.7 Ma. During this period, and based on the present day outcropping sequences, the mean crustal growth rate varied from 32 to 65 km(3)/km/Ma (volume per unit width along the strike of the arc) which is comparable to the range for present day arcs of the Western Pacific region. The correspondence of crustal growth rates observed for present-day island arcs and the good preservation of the crustal section in the Kohistan arc, make this area an exceptional natural laboratory to study arc related processes and to check models of continental crust formation by arc accretion.
ISSN: 0012-821X

hal-00644555
https://hal.archives-ouvertes.fr/hal-00644555
DOI : 10.1016/j.epsl.2011.07.016

Éditeur(s) : Pêche thonière et dispositifs de concentration de poissons, Caribbean-Martinique, 15-19 Oct 1999
Résumé : This paper develops a comparative overview of the development of Fish Aggregating Devices (or fads) by the tropical purse seine fisheries in various ocean areas (Atlantic, Indian, Western and Eastern Pacific Oceans). First, a comparative review of fishery statistics is developed. Recent catches on fads by purse seiners can be estimated at a level of one million tons yearly. This comparison allows to describe the fishing zones and catch trends over time, the species composition and size taken in the various fad fisheries. Skipjack tuna is the dominant species in most fad fisheries, but significant amounts of yellowfin and bigeye are also frequently taken under fads. The paper also presents an overview of various observer data concerning the by-catches of the various fisheries using fads. This comparison allows to estimate the total worldwide yearly by-catches at about 100,000 t, and their species composition in each ocean. The fad technology used in the various oceans by purse seiners is described. The recent use of fads by purse seiners has introduced major uncertainties in most stock assessments, because analyses are hampered by changes in the fishing effort in a fad fishery, by changes in fishing zones and in sizes caught. As a last point, the management of fads presently done or in view by the various tuna bodies are introduced. It appears that the present massive use of fads worldwide is perhaps an unsafe fishing mode, which could produce serious overfishing of many stocks. There is then a consensus that the use of fads needs to be controlled and limited to sustainable biological levels.

Droits : Ifremer
http://archimer.ifremer.fr/doc/00042/15278/12664.pdf
http://archimer.ifremer.fr/doc/00042/15278/

Éditeur(s) : Gauthier-Villars
Résumé : Analysis of Geosat sea level data over the period November 1986-November 1987 has evidenced long, low-frequency equatorial wave propagations over most of the equatorial Pacific basin (Delcroix et al.,1991). The succession of downwelling Kelvin, upwelling Kelvin, and upwelling Rossby waves has raised a series of questions. Is the observed upwelling Rossby wave due to a reflection process on the eastern boundary, or due to wind forcing near the equator? Why is there no evidence of a reflection of the downwelling Kelvin wave? Is this wave sequence a feature of the 1986-1987 El Nino phenomenon or is it to be found in the mean seasonal cycle? We used a linear model forced by the FSU wind stress to study these questions. The results, during the period November 1986-November 1987, show the successive downwelling/ upwelling Kelvin waves and upwelling Rossby wave as in the Geosat observations. Another test was made, using only the part of the forcing responsible for creating Kelvin waves, in order to evaluate the portion of the signals that was due to the reflection process and the part due to local forcing. It appears that wind forcing favourable to upwelling in the eastern part of the equatorial basin is the main mechanism which both generates the upwelling Rossby wave and inhibits the reflection of the downwelling Kelvin wave. However, the reflection of the upwelling Kelvin wave contributes to the upwelling Rossby wave signal, and makes it visible all the way from the eastern to the western sides of the equatorial Pacific. Results of a climatological run show that the successive downwelling/upwelling Kelvin and Rossby wave propagations are present in the mean seasonal cycle. However, the corresponding model signals, as well as the wind forcing anomalies, show a larger amplitude in 1986-1987.
Oceanologica Acta (0399-1784) (Gauthier-Villars), 1992 , Vol. 15 , N. 5 , P. 545-554

Droits : info:eu-repo/semantics/openAccess
http://archimer.ifremer.fr/doc/00101/21180/18797.pdf
http://archimer.ifremer.fr/doc/00101/21180/

Éditeur(s) : HAL CCSD Elsevier
Résumé : We present new analytical data from lavas and associated rocks dredged and/or drilled from the South Fiji Basin, Northland Plateau, Colville Ridge and Havre Trough. These results provide much-needed ground truth about the geology, age and tectonic evolution of the Cenozoic submarine ridges and basins between the active intraoceanic Tonga–Kermadec arc, and rifted continental borderlands of New Zealand, the Norfolk Ridge and New Caledonia. Key results from this study include: (1) Ar–Ar dates on Minerva Abyssal Plain oceanic crust suggest that the ages of magnetic anomalies in the South Fiji Basin have been overestimated by earlier workers; (2) subduction-related lavas are widespread across the region, are not presently organised into arc-like chains, and cluster in the age range 22–18 Ma (Early Miocene); (3) the oldest subduction-related lavas occur in the western part of the region (32–26 Ma: Norfolk and Three Kings Ridge); (4) shoshonites, interpreted as rifted arc lavas, were erupted in a narrow 20–21 Ma interval over a wide area. Put together, these results indicate high magmatic flux and large and rapid horizontal tectonic translations and basin opening from 18–23 Ma in the region immediately north of New Zealand. We explain the Miocene tectonomagmatic development of the region by a model of rapid rollback of a single, east-facing Pacific arc–trench system that became established after Northland Allochthon emplacement. Critical testing of this, versus other, tectonic models must await drilling and dating of thus-far unsampled Kupe Abyssal Plain crust.
ISSN: 0025-3227

hal-00405803
https://hal.archives-ouvertes.fr/hal-00405803
DOI : 10.1016/j.margeo.2006.10.033

Éditeur(s) : La recherche face aux pêches artisanales
Résumé : Each of nine people agreed to write a summary regarding changes in research on small-scale fishing in a large region of the world. These summaries that will be presented to you focus respectively on changes in research in Southeast Asia, in the South Pacific along the western coast of Latin America, in West Africa, in the Mediterranean, in Northwest Europe, and in North America. In order to help you reconstruct their actions in the halieutic context of the regions about which they are talking, Serge Garcia will first present us with a panorama of small-scale fishing. By asking them to do this study, we wished to be able to place the symposium's study within an overall framework and in an historical perspective. The objective was to relativise the conclusions that could be drawn from analyses of local situations, or, on the contrary, to allow them to be generalised by using bases of comparison. The summaries are far from being able to offer a planetary geographical coverage, just as they do not really include the evolution of all disciplines. The means available to the organisers did not allow for going up to that point. We must at this point thank the authors who agreed to do this complex work of summarisation, because they succeeded in showing the richness of such a step, which the participants will be able to appreciate in their reports. In this analysis of geographical syntheses, we would like to draw a few comparisons able to fuel the symposium's debates. We will examine how small-scale fishing became a subject of research, and the difficulty of defining it. Then we will see how the various disciplines became mutually involved, then specialised, depending upon the stages through which the research passed on this subject. The role of institutions and the weight of the contextual environment will be examined. Finally, we will propose a few possible directions for the debates on various themes of the symposium. Some outside contributions to the summaries also focus on the evolution of the research and we have taken that into account in this presentation. Limited to one country (Yahaya Jahara) or to one province (Maneschy and Furtado), they are useful complements to the summaries.
(OCR non controlé) Neuf personnes ont accepté de rédiger chacune une synthèse concernant l'évolution des recherches sur les pêches artisanales dans une grande région du monde. Les synthèses qui vont vous être présentées portent respectivement sur l'évolution des recherches en Asie du Sud-Est, dans le Pacifique-sud sur la façade occidentale de l'Amérique Latine, en ,Afrique de l'Ouest, en Méditerranée, en Europe du Nord- Ouest, en Amérique du Nord. Pour vous permettre de resituer leurs interventions dans le contexte halieutique des régions dont ils nous parlent, Serge Garcia nous présentera en premier lieu un panorama des pêches artisanales. En leur demandant ce travail, nous avons souhaité pouvoir replacer les travaux du symposium dans un cadre global et dans une perspective historique. L'objectif était de relativiser les conclusions qui pourraient être tirées d'analyses de situations locales, ou, au contraire, de permettre leur généralisation en disposant de bases de comparaison. Les synthèses sont loin d'offrir une couverture géographique planétaire, tout comme elles n'intègrent pas vraiment l'évolution de toutes les disciplines. Les moyens à la disposition des organisateurs ne permettaient pas d'aller jusque là. Il nous faut remercier ici les auteurs qui ont consenti à faire ce travail complexe de synthèse, car ils ont réussi à démontrer la richesse d'une telle démarche, ainsi que les participants pourront l'apprécier à travers leurs exposés. Dans cette analyse des synthèses géographiques, nous voudrions tenter quelques comparaisons de nature à alimenter les débats du symposium. Nous examinerons comment est née la pêche artisanale en tant qu'objet de recherche, et la difficulté de la définir. Puis nous verrons, en fonction des étapes par lesquelles sont passées les recherches sur ce sujet, comment les différentes disciplines se sont mutuellement impliquées, puis se sont spécialisées. Le rôle des institutions et le poids des contextes seront examinés. Enfin nous proposerons quelques orientations possibles pour les débats sur les différents thèmes du symposium. Certaines contributions extérieures aux synthèses portent aussi sur l'évolution des recherches et nous en avons tenu compte dans cette présentation. Limitées à un pays (Yahaya Jahara) ou à une province (Maneschy et Furtado), elles complètent utilement les synthèses.

Droits : info:eu-repo/semantics/openAccess
http://archimer.ifremer.fr/doc/1990/acte-2464.pdf
http://archimer.ifremer.fr/doc/00000/2464/

Éditeur(s) : Springer
Résumé : Pearl oysters belonging to the genus Pinctada (Bivalvia: Pteriidae) are widely distributed between the Indo-Pacific and western Atlantic. The existence of both widely distributed and more restricted species makes this group a suitable model to study diversification patterns and prevailing modes of speciation. Phylogenies of eight out of the 11 currently recognised Pinctada species using mitochondrial (cox1) and nuclear (18S rRNA) data yielded two monophyletic groups that correspond to shell size and presence/absence of hinge teeth. Character trace of these morphological characters onto the molecular phylogeny revealed a strong correlation. Pinctada margaritifera appears polyphyletic with specimens from Mauritius grouping in a different clade from others of the French Polynesia and Japan. Hence, P. margaritifera might represent a species complex, and specimens from Mauritius could represent a different species. Regarding the putative species complex Pinctada fucata/Pinctada martensii/Pinctada radiata/Pinctada imbricata, our molecular analyses question the taxonomic validity of the morphological characters used to discriminate P. fucata and P. martensii that exhibited the lowest genetic divergence and are most likely conspecific as they clustered together. P. radiata and P. imbricata were recovered as monophyletic. The absence of overlapping distributions between sister lineages and the observed isolation by distance suggests that allopatry is the prevailing speciation mode in Pinctada. Bayesian dating analysis indicated a Miocene origin for the genus, which is consistent with the fossil record. The northward movement of the Australian plate throughout the Miocene played an important role in the diversification process within Pinctada.
Marine Biotechnology (1436-2228) (Springer), 2011-04 , Vol. 13 , N. 2 , P. 181-192

Droits : Springer Science+Business Media, LLC 2010
http://archimer.ifremer.fr/doc/00035/14635/14015.pdf
DOI:10.1007/s10126-010-9278-y
http://archimer.ifremer.fr/doc/00035/14635/

Éditeur(s) : Springer
Résumé : Series of sensitivity tests were performed with a z-coordinate, global eddy-permitting (1/4 degrees) ocean/sea-ice model (the ORCA-R025 model configuration developed for the DRAKKAR project) to carefully evaluate the impact of recent state-of-the-art numerical schemes on model solutions. The combination of an energy-enstrophy conserving (EEN) scheme for momentum advection with a partial step (PS) representation of the bottom topography yields significant improvements in the mean circulation. Well known biases in the representation of western boundary currents, such as in the Atlantic the detachment of the Gulf Stream, the path of the North Atlantic Current, the location of the Confluence, and the strength of the Zapiola Eddy in the south Atlantic, are partly corrected. Similar improvements are found in the Pacific, Indian, and Southern Oceans, and characteristics of the mean flow are generally much closer to observations. Comparisons with other state-of-the-art models show that the ORCA-R025 configuration generally performs better at similar resolution. In addition, the model solution is often comparable to solutions obtained at 1/6 or 1/10 degrees resolution in some aspects concerning mean flow patterns and distribution of eddy kinetic energy. Although the reasons for these improvements are not analyzed in detail in this paper, evidence is shown that the combination of EEN with PS reduces numerical noise near the bottom, which is likely to affect current-topography interactions in a systematic way. We conclude that significant corrections of the mean biases presently seen in general circulation model solutions at eddy-permitting resolution can still be expected from the development of numerical methods, which represent an alternative to increasing resolution.
Ocean Dynamics (1616-7341) (Springer), 2006-12 , Vol. 56 , N. 5-6 , P. 543-567

Droits : © Springer 2006. Part of Springer Science+Business Media
http://archimer.ifremer.fr/doc/2006/publication-3514.pdf
DOI:10.1007/s10236-006-0082-1
http://archimer.ifremer.fr/doc/00000/3514/

Éditeur(s) : Wiley / Blackwell
Résumé : Six cohorts of the silver-stripe round herring Spratelloides gracilis, a fast-growing and short-lived tropical clupeid, were collected as juveniles and then as adults during austral summers from November to February in 1998–1999 and 1999–2000, using light traps in the Dampier Archipelago, Western Australia. Otolith analysis allowed backcalculation of size and growth rate at age to examine the relative influences of selective mortality and water temperature on early growth. Negative sizeselective mortality and growth-selective mortality between the juvenile and the adult stages was found only in the cohort that was the smallest and slowest growing in the period immediately following hatching. Selective mortality preferentially removed members of this cohort that were smaller from age 0 to 15 days, and slower growing from 0 to 10 days, resulting in an elevation of size at age to, or even above, that of cohorts that had not undergone this process. Size and growth rate at 5 day age intervals in the first 20 days after hatching differed among cohorts within and between summers and were strongly and positively correlated (r2= 0·61–0·83) with water temperature.
Journal of Fish Biology (0022-1112) (Wiley / Blackwell), 2009-03 , Vol. 74 , N. 4 , P. 921-938

Droits : 2010 The Fisheries Society of the British Isles, Wiley Blackwell Publishing, Inc.
http://archimer.ifremer.fr/doc/00008/11940/8643.pdf
DOI:10.1111/j.1095-8649.2009.02179.x
http://archimer.ifremer.fr/doc/00008/11940/