Éditeur(s) : HAL CCSD Wiley-Blackwell
Résumé :  
Cross-species analyses of plant functional traits have shed light on factors contributing to differences in performance and distribution, but to date most studies have focused on either leaves or stems. We extend these tissue-specific analyses of functional strategy towards a whole-plant approach by integrating data on functional traits for 13 448 leaves and wood tissues from 4672 trees representing 668 species of Neotropical trees. Strong correlations amongst traits previously defined as the leaf economics spectrum reflect a tradeoff between investments in productive leaves with rapid turnover vs. costly physical leaf structure with a long revenue stream. A second axis of variation, the 'stem economics spectrum', defines a similar tradeoff at the stem level: dense wood vs. high wood water content and thick bark. Most importantly, these two axes are orthogonal, suggesting that tradeoffs operate independently at the leaf and at the stem levels. By simplifying the multivariate ecological strategies of tropical trees into positions along these two spectra, our results provide a basis to improve global vegetation models predicting responses of tropical forests to global change.
Ecology Letters

hal-01032142
https://hal.archives-ouvertes.fr/hal-01032142
DOI : 10.1111/j.1461-0248.2010.01517.x

Éditeur(s) : HAL CCSD
Résumé :  
In tropical rainforests, the origin of the interspecific variability in trunk CO2 efflux (Et) and of seasonal variations in Et have not yet been explained. We conducted a study to test which biotic and abiotic determinants would contribute to these variations. Et and a set of wood anatomical and functional characteristics were measured on trees in two habitats (Terra Firme forest vs. seasonal flooded forest) in a Neotropical rainforest over a 18-months' period including a range of climatic and soil water conditions. The large variability in Et (0.33 to 3.15 μmol CO2 m-2 s-1 in Terra Firme forest) was associated with tree diameter and growth respiration (Eg), although maintenance respiration (Em) was the main component of Et. Neither wood properties nor sapflow density were correlated with trunk CO2 efflux. The large seasonal variations in Et were correlated with diameter growth and surface soil water content (SWC). These data underlined the absence of clearly identified biotic determinants, beside growth, to explain the interspecific variability in Et. We emphasized the fast response of growth and maintenance respiration processes to seasonal variations in SWC that potentially has consequences on the carbon cycle in the ecosystem under upcoming climatic changes.
23. New Phytologist Symposium. Carbon cycling in tropical ecosystems
Guangzhou, China

hal-01032199
https://hal.archives-ouvertes.fr/hal-01032199

Éditeur(s) : HAL CCSD
Résumé :   Diplôme : DEA
il s'agit d'un type de produit dont les métadonnées ne correspondent pas aux métadonnées attendues dans les autres types de produit : DISSERTATION
Tropical rainforests are complex ecosystems, rarely investigated from ground-level. The canopy, highest part of the cover, can on the other hand be studied by several remote sensing tools. The aim of this study is to assess in what way the canopy is related to the forest structure below and the forest dynamic. We looked for correlation between the canopy texture (crown size) observed from aerial photographs and various parameters of the forest (spatial and demographic structure, forest dynamic, floristic structure). We worked on the Paracou experimental device, in French Guiana, at two different scales. 1- At a local scale (6.5 ha ; scale of the photographs 1: 4 000), we demarcated six sub-plots with big crowns in the canopy and six with small crowns. 2- At a forest scale (approximately one hundred ha ; scale of the photograph 1: 25 000), we compared 2 forest plots: one with a rough canopy texture and one with a smooth canopy texture. We found that the size of the crowns in the canopy do not express the same characteristics of the fore according to the scale of study : at the local scale, difference in crown size means a difference in basal area but not in density, while at the forest scale it is the opposite. Several hypotheses related to the sylvigenetic cycle, the floristic structure or the nature of the substratum are suggested to explain these results.
Les forêts tropicales humides (FTH) sont des écosystèmes complexes difficiles à étudier depuis le sol. Leur canopée, partie supérieure du couvert, est en revanche abordable par divers outils de télédétection. Cette étude vise à déterminer en quoi la structure de la canopée est le reflet de la structure forestière sous-jacente et de la dynamique spatio-temporelle qui s'y exerce. Nous avons donc recherché des corrélations, entre la texture de canopée (taille des couronnes) vue sur des photographies aériennes, et différents paramètres de la forêt sous-jacente (structure spatiale et démographique, dynamique forestière, structure floristique). Ce travail a été abordé sur le dispositif expérimental forestier de Paracou, en Guyane Française, à 2 échelles différentes. 1- A l'échelle de la parcelle (6,5 ha ; photographies au 1: 4 000), nous avons délimité des placettes à grosses couronnes en canopée et des placettes à petites couronnes. 2- A l'échelle du massif forestier (une centaine d'ha ; photographies IGN au 1: 25 000) nous comparons une parcelle de texture de canopée, grossière et une parcelle de texture fine. Le résultat principal de notre étude est que la taille des couronnes en canopée ne reflète pas les mêmes caractéristiques du couvert selon l'échelle de l’étude : à l'échelle de la parcelle, des différences de taille de couronnes traduisent des différences de surface terrière mais pas de densité, tandis qu'à l'échelle du massif forestier c'est l'inverse. Plusieurs hypothèses, invoquant des différences dans le stade sylvigénétique, la composition floristique ou la nature du substrat des placettes ou parcelles comparées, sont proposées pour expliquer ces résultats.
https://hal.archives-ouvertes.fr/hal-01189180

hal-01189180
https://hal.archives-ouvertes.fr/hal-01189180
PRODINRA : 12184

Éditeur(s) : HAL CCSD American Geophysical Union (AGU)
Résumé : International audience
Nowadays, technological advances in satellite imagery measurements as well as the development of dense geodetic and seismologic networks allow for a detailed analysis of surface deformation associated with active fault seismic cycle. However, the study of earthquake dynamics faces several limiting factors related to the difficulty to access the deep source of earthquake and to integrate the characteristic time scales of deformation processes that extend from seconds to thousands of years. To overcome part of these limitations and better constrain the role and couplings between kinematic and mechanical parameters, we have developed a new experimental approach allowing for the simulation of strike-slip fault earthquakes and analyze in detail hundreds of successive seismic cycle. Model rheology is made of multilayered visco-elasto-plastic analog materials to account for the mechanical behavior of the upper and lower crust and to allow simulating brittle/ductile coupling, postseismic deformation phase and far-field stress transfers. The kinematic evolution of the model surface is monitored using an optical system, based on subpixel spectral correlation of high-resolution digital images. First, results show that the model succeed in reproducing the deformation mechanisms and surface kinematics associated to the main phases of the seismic cycle indicating that model scaling is satisfactory. These results are comforted by using numerical algorithms to study the strain and stress distribution at the surface and at depth, along the fault plane. Our analog modeling approach appears, then, as an efficient complementary approach to investigate earthquake dynamics.
ISSN: 0278-7407

hal-01174176
https://hal.archives-ouvertes.fr/hal-01174176
DOI : 10.1002/2014TC003701

Éditeur(s) : HAL CCSD Oxford University Press (OUP)
Résumé : International audience
A temporary network of 10 broad-band seismic stations has been installed in French Polynesia for the Polynesian Lithosphere and Upper Mantle Experiment (PLUME). All the seismic stations were installed either on volcanic islands or on atolls of the various archipelagos of French Polynesia in a manner which complements the geographic coverage provided by the regional permanent stations. The primary aim of PLUME is to image the upper mantle structures related to plate motion and hotspot activity. However, because of its proximity to all sites, the ocean is responsible for a high level of noise in the seismic data and we show that these data can also be used to analyse ocean wave activity. The power spectral density (PSD) analyses of the seismic data recorded in French Polynesia show clear peaks in the 0.05– 0.10 Hz band (periods between 10 and 20 s), which corresponds to swell frequencies. Clear peaks in this frequency band are also observed in infrasonic data recorded on Tahiti. Ground motion analysis shows that the swell-related seismic noise (SRSN) is linearly polarized in the horizontal plane and its amplitude decreases rapidly with the distance from the shore. The microseismic and the infrasonic 'noise' amplitudes show very similar variations from station to station and both are strongly correlated with the swell amplitudes predicted by the National Oceanic and Atmospheric Administration (NOAA), wind-forced, 'WaveWatch' models. The swell direction can be estimated from SRSN polarization analysis but this has to be done with care since, for some cases, the ground motions are strongly controlled by the islands' anisometric shapes and by swell refraction processes. We find cases, however, such as Tahiti or roughly circular Tuamotu atolls, where the azimuth of the swell is in good agreement with the seismic estimates. We, therefore, demonstrate that the SRSN and the infrasonic signal observed in French Polynesia can be used in such cases as a proxy for swell amplitude and azimuth. From the continuous analysis of the data recorded in 2003 at the permanent seismic station PPTL in Tahiti, transfer functions have been obtained. This could provide a way to quantify the swell activity during the last two decades and, therefore, assist in the investigation of climate changes.
ISSN: 0956-540X

hal-01388809
http://hal.univ-reunion.fr/hal-01388809
http://hal.univ-reunion.fr/hal-01388809/document
http://hal.univ-reunion.fr/hal-01388809/file/Barruol_swell_GJI2006.pdf
DOI : 10.1111/j.1365-246X.2006.02871.x

Éditeur(s) : HAL CCSD
Résumé : International audience
The most active deep convection area in the western Mediterranean Sea is located in the Gulf of Lions. Recent studies in this area provides some insights on the complexity of the physical dynamics of convective regions, but very little is known about their impacts on the biogeochemical properties. The CASCADE (CAscading, Surge, Convection, Advection and Downwelling Events) cruise, planed in winter 2011, give us the opportunity to compare vertical profiles of properties sampled either during stratified conditions or after/during a convection event. In the present study, we focus on the distributions of the carbonate system properties (mainly total alkalinity, AT; and total dissolved inorganic carbon, CT) because, in the context of the climate change, deep convection areas are suspected to significantly increase the sequestration of anthropogenic CO2 (CANT). Given its limited size, the impact of the Mediterranean Sea on the global carbon budget is probably minor but this marginal sea can be used as a laboratory to better understand carbon sequestration and its transfer to the basin interior by deep convection processes. Distributions of AT and CT, both measured from bottle samples, and that of CANT (estimated with the TrOCA approach) are first analyzed in the light of other key properties (salinity, temperature, and dissolved oxygen). An objective interpolation procedure is then applied to estimate CT and AT from CTD measured properties. With this procedure, the vertical resolution goes from a maximum of 32 samples per station to one property estimate every meter (more detailed distributions are obtained). Results provide arguments to conclude that CANT is rapidly transferred to the deepest layer due to deep convection events. During deep convection events, the increase of CANT in the water column is positively correlated to that of potential density and oxygen content. The challenge of quantifying the amount of sequestered carbon is however not resolved due to the complexity and the highly dynamical nature of the convective regions. Deep convection in the Gulf of Lions, in parallel with cascading along the continental slope, could thus potentially explain the very high levels of both CANT and acidification estimated in the deep layers of the western Mediterranean Sea.
Deep Sea Research Part I: Oceanographic Research Papers

hal-01336725
https://hal-univ-perp.archives-ouvertes.fr/hal-01336725
DOI : 10.1016/j.dsr.2016.04.003

Éditeur(s) : HAL CCSD Wiley
Résumé : International audience
1. Despite considerable interest in the application of plant functional traits to questions of community assembly and ecosystem structure and function, there is no consensus on the appropriateness of sampling designs to obtain plot-level estimates in diverse plant communities. 2. We measured 10 plant functional traits describing leaf and stem morphology and ecophysiology for all trees in nine 1-ha plots in terra firme lowland tropical rain forests of French Guiana (N = 4709). 3.We calculated, by simulation, the mean and variance in trait values for each plot and each trait expected under seven sampling methods and a range of sampling intensities. Simulated sampling methods included a variety of spatial designs, as well as the application of existing data base values to all individuals of a given species. 4. For each trait in each plot, we defined a performance index for each sampling design as the proportion of resampling events that resulted in observed means within 5% of the true plot mean, and observed variance within 20%of the true plot variance. 5. The relative performance of sampling designs was consistent for estimations of means and variances. Data base use had consistently poor performance for most traits across all plots, whereas sampling one individual per species per plot resulted in relatively high performance. We found few differences among different spatial sampling strategies; however, for a given strategy, increased intensity of sampling resulted in markedly improved accuracy in estimates of trait mean and variance. 6. We also calculated the financial cost of each sampling design based on data from our 'every individual per plot' strategy and estimated the sampling and botanical effort required. The relative performance of designs was strongly positively correlated with relative financial cost, suggesting that sampling investment returns are relatively constant. 7. Our results suggest that trait sampling for many objectives in species-rich plant communities may require the considerable effort of sampling at least one individual of each species in each plot, and that investment in complete sampling, though great, may be worthwhile for at least some traits.
ISSN: 0269-8463

hal-00594009
https://hal.archives-ouvertes.fr/hal-00594009

Éditeur(s) : HAL CCSD Geological Society of London
Résumé : International audience
Low-velocity zones ('bright spots') imaged by the INDEPTH seismic experiment in southern Tibet are extensively interpreted as widespread partial melt within the crust, which has given a strong support for the channel flow model. These suggest that a continuous seismic low-velocity zone underlies Tibet on the large scale. Here we take advantage of the Hi-CLIMB seismic experiment which includes a dense south-north profile and a lateral 2D seismic network to assess the vertical and the horizontal extension of low-velocity zones in southern Tibet. Several approaches including migration, amplitude analysis and waveform inversion of receiver functions are performed to detect crustal low-velocity zones using this new seismological dataset. Our results reveal localized and discontinuous low-velocity zones in Tibet. They indicate that the vertical extension of the low-velocity zones is about 10 km, and their maximum horizontal length appears to be c. 50 km. Our study suggests a partial correlation between the location of these low-velocity zones and the spatial distribution of Tibetan grabens. These results, especially the non-continuity of low-velocity zones, together with the observed regular value of mean crustal VP/VS ratio, question the existence of widespread partial melt of the southern Tibetan crust and, therefore, the viability of the channel flow model.
Growth and Collapse of the Tibetan Plateau

hal-00853971
https://hal.archives-ouvertes.fr/hal-00853971
DOI : 10.1144/SP353.6

Éditeur(s) : HAL CCSD Oxford University Press (OUP)
Résumé : International audience
A temporary network of 10 broad-band seismic stations has been installed in French Polynesia for the Polynesian Lithosphere and Upper Mantle Experiment (PLUME). All the seismic stations were installed either on volcanic islands or on atolls of the various archipelagos of French Polynesia in a manner which complements the geographic coverage provided by the regional permanent stations. The primary aim of PLUME is to image the upper mantle structures related to plate motion and hotspot activity. However, because of its proximity to all sites, the ocean is responsible for a high level of noise in the seismic data and we show that these data can also be used to analyse ocean wave activity. The power spectral density (PSD) analyses of the seismic data recorded in French Polynesia show clear peaks in the 0.05– 0.10 Hz band (periods between 10 and 20 s), which corresponds to swell frequencies. Clear peaks in this frequency band are also observed in infrasonic data recorded on Tahiti. Ground motion analysis shows that the swell-related seismic noise (SRSN) is linearly polarized in the horizontal plane and its amplitude decreases rapidly with the distance from the shore. The microseismic and the infrasonic 'noise' amplitudes show very similar variations from station to station and both are strongly correlated with the swell amplitudes predicted by the National Oceanic and Atmospheric Administration (NOAA), wind-forced, 'WaveWatch' models. The swell direction can be estimated from SRSN polarization analysis but this has to be done with care since, for some cases, the ground motions are strongly controlled by the islands' anisometric shapes and by swell refraction processes. We find cases, however, such as Tahiti or roughly circular Tuamotu atolls, where the azimuth of the swell is in good agreement with the seismic estimates. We, therefore, demonstrate that the SRSN and the infrasonic signal observed in French Polynesia can be used in such cases as a proxy for swell amplitude and azimuth. From the continuous analysis of the data recorded in 2003 at the permanent seismic station PPTL in Tahiti, transfer functions have been obtained. This could provide a way to quantify the swell activity during the last two decades and, therefore, assist in the investigation of climate changes.
ISSN: 0956-540X

hal-01323691
http://hal.univ-reunion.fr/hal-01323691
http://hal.univ-reunion.fr/hal-01323691/document
http://hal.univ-reunion.fr/hal-01323691/file/Barruol_et_al_2006.pdf
DOI : 10.1111/j.1365-246X.2006.02871.x

Éditeur(s) : HAL CCSD Montpellier : CIRAD
Résumé : This article presents the effects of logging on the dynamics of above-ground biomass from the results of the post-logging study within two forests: Cikel in Eastern Para, Brazil and Paracou in French Guiana. The main objective is to compare the impact of commercial logging on the regeneration of the above-ground biomass in these forests whose characteristics differ in terms of structure and growth. In both sites, the intensity of exploitation is a key factor in determining the loss of biomass and the time required for its regeneration. In Paracou, the regeneration of biomass lost during conventional logging of 10 trees per hectare takes 45 years and more than 100 years when operating with higher intensity (21 trees/ha). In Cikei the forest biomass regenerates after 49 years harvesting 6 trees/ha and that takes 87 years after removal of 8 trees/ha. This regeneration needs similar time on both sites but with lower logging intensity at Cikel, in which felled trees are larger with a greater biomass than those of Paracou. This post-logging study has established a direct correlation of the dynamics of the biomass with the initial structure of the forest, as well as with the parameters of forest dynamics: mortality, growth and recruitment. The accumulation of biomass by the tree growth of the two remaining stands is a key parameter for the net carbon storage, while the contribution of recruitment in Paracou becomes significant only after 10 years after felling. Therefore in view to improve the growth of residual trees, it is compulsory to apply adequate silvicultural treatments such as selective thinning or removal of vines. While the two forests are geographically close enough, their regenerative abilities differ and because of the significant difference in size of the trees, the forest could tolerate more intensive harvesting in French Guiana
ISSN: 0006-579X

hal-01032427
https://hal.archives-ouvertes.fr/hal-01032427

Éditeur(s) : HAL CCSD Springer Verlag
Résumé : International audience
In this study, we quantify and analyze the spatial and temporal evolution of the surface displacement of Kahrod landslide located in the center of Alborz range (Iran) within the Haraz valley. This landslide represents a threat for this main drainage axis and its numerous infrastructures. We present three sets of displacement vectors based on GPS technique. An 8-benchmark network has been surveyed four times on a 1-year period basis. It provides accurate information on the rate of displacement within the landslide, and addresses the problem of the mechanical resistance of a small hillock, down slope, under the stress imposed by the landslide. Then. this network is densified (57 marks) and measured twice in 6 months using a rapidstatic approach. This yields to a dense description of surface deformation over the whole landslide. Finally, a 1-year time series of permanent GPS recordings is presented and compared to rainfall. Furthermore, we analyze Envisat radar differential interferograms (DInSAR) spanning the same period as permanent GPS. These geodetic data allow to precisely determine the limits of the current sliding zone and to describe the spatial and temporal evolution of surface displacement. The combination of geodesy and field observations leads to a precise description of the past and present kinematics behavior of Kahrod landslide. The chaotic nature of the sliding mass suggests a first catastrophic landslide in a first episode. During the period of observation, the landslide appears to deform quite steadily, and the evidence of short-term correlation between rainfall and deformation amplitude needs to be confirmed by future measurements. Carrying on the acquisition of GPS and InSAR data within the sliding mass but also within adjacent bedrock should give fundamental information with regards to major activation processes (river sapping, water seeping, earthquakes, or failure within the frontal hill of bedrock) and their potential consequences.
ISSN: 1435-9529

hal-00411318
https://hal.archives-ouvertes.fr/hal-00411318
DOI : 10.1016/j.enggeo.2008.02.013

Éditeur(s) : HAL CCSD American Geophysical Union
Résumé : International audience
Analysis of continuous seismic data recorded by a dense passive seismological network (Hi-CLIMB) installed across the Himalayas reveals strong spatial and temporal variations in the ambient seismic energy produced at high frequencies (>1 Hz). From June to September 2003, the high-frequency seismic noise is observed to increase up to 20 dB (relative to (m/s)(2)/Hz) for all the stations located along a steep 30-km-long narrow and deeply incised channel of the Trisuli River, a major trans-Himalayan river. The early summer increase in high-frequency energy is modulated by a 24-h periodicity where the minimum of seismic noise level is reached around noon and the maximum is reached late in the evening. A detailed study of seismic noise amplitude reveals a clear correlation with both regional meteorological and hydrological data along the Trisuli River. Seasonal increase in ambient noise coincides with the strong monsoon rainfall and a period of rapid melting of snow and ice in the high elevations. The observed 24-h cyclicity is consistent with the daily fluctuation of the precipitation and river discharge in the region. River-induced seismic noise is partly generated by stream turbulence, but this mechanism fails to explain the observed clockwise hysteresis of seismic noise amplitude versus water level. This pattern is better explained if a significant part of the observed seismic noise is caused by ground vibrations generated by bed load transport. This points out the potential of using background seismic noise to quantify in continuous river bed load and monitor its spatial variations, which remain difficult with classical approaches.
ISSN: 2169-9313

hal-00412170
https://hal.archives-ouvertes.fr/hal-00412170
DOI : 10.1029/2007JB005034

Éditeur(s) : HAL CCSD Wiley-Blackwell
Résumé :  
1. Despite considerable interest in the application of plant functional traits to questions of community assembly and ecosystem structure and function, there is no consensus on the appropriateness of sampling designs to obtain plot-level estimates in diverse plant communities. 2. We measured 10 plant functional traits describing leaf and stem morphology and ecophysiology for all trees in nine 1-ha plots in terra firme lowland tropical rain forests of French Guiana (N = 4709). 3.We calculated, by simulation, the mean and variance in trait values for each plot and each trait expected under seven sampling methods and a range of sampling intensities. Simulated sampling methods included a variety of spatial designs, as well as the application of existing data base values to all individuals of a given species. 4. For each trait in each plot, we defined a performance index for each sampling design as the proportion of resampling events that resulted in observed means within 5% of the true plot mean, and observed variance within 20%of the true plot variance. 5. The relative performance of sampling designs was consistent for estimations of means and variances. Data base use had consistently poor performance for most traits across all plots, whereas sampling one individual per species per plot resulted in relatively high performance. We found few differences among different spatial sampling strategies; however, for a given strategy, increased intensity of sampling resulted in markedly improved accuracy in estimates of trait mean and variance. 6. We also calculated the financial cost of each sampling design based on data from our 'every individual per plot' strategy and estimated the sampling and botanical effort required. The relative performance of designs was strongly positively correlated with relative financial cost, suggesting that sampling investment returns are relatively constant. 7. Our results suggest that trait sampling for many objectives in species-rich plant communities may require the considerable effort of sampling at least one individual of each species in each plot, and that investment in complete sampling, though great, may be worthwhile for at least some traits.
Functional Ecology

hal-01032175
https://hal.archives-ouvertes.fr/hal-01032175
DOI : 10.1111/j.1365-2435.2009.01600.x

Éditeur(s) : HAL CCSD
Résumé : International audience
Extreme rainfall events can trigger numerous landslides in mountainous areas and a prolonged increase of river sediment load. The resulting mass transfer at the Earth surface in turn induces stress changes at depth, which could be sufficient to trigger shallow earthquakes. The 2009 Morakot typhoon represents a good case study as it delivered 3 m of precipitation in 3 days and caused some of the most intense erosion ever recorded. Analysis of seismicity time-series before and after the Morakot typhoon reveals a systematic increase of shallow (i.e. 0-15 km of depth) earthquake frequency in the vicinity of the areas displaying a high spatial density of landslides. This step-like increase in frequency lasts for at least 2-3 years and does not follow an Omori-type aftershock sequence. Rather, it is associated to a step change of the Gutenberg-Richter b-value of the earthquake catalog. Both changes occurred in mountainous areas of southwest Taiwan, where typhoon Morakot caused extensive landsliding. These spatial and temporal correlations strongly suggest a causal relationship between the Morakot-triggered landslides and the increase of earthquake frequency and their associated b-value. We propose that the progressive removal of landslide materials from the steep mountain landscape by river sediment transport acts as an approximately constant increase of the stress rate with respect to pre-typhoon conditions, and that this in turn causes a step-wise increase in earthquake frequency. To test this hypothesis, we investigate the response of a rate-and-state fault to stress changes using a 2-D continuum elasto-dynamic model. Consistent with the results of Ader et al. (2013), our preliminary results show a step-like increase of earthquake frequency in response to a step-like decrease of the fault normal stress. We also investigate the sensitivity of the amplitude and time-scale of the earthquake frequency increase to the amplitude of the normal stress change and to rheological parameters. Our study offers new insights on the potential influence of extreme erosional events on the short-time scale dynamics of faults and earthquakes.
American Geophysical Union Fall Meeting 2016
San Francisco, United States

insu-01416987
https://hal-insu.archives-ouvertes.fr/insu-01416987