Éditeur(s) : HAL CCSD
Résumé : International audience
Extreme rainfall events can trigger numerous landslides in mountainous areas and a prolonged increase of river sediment load. The resulting mass transfer at the Earth surface in turn induces stress changes at depth, which could be sufficient to trigger shallow earthquakes. The 2009 Morakot typhoon represents a good case study as it delivered 3 m of precipitation in 3 days and caused some of the most intense erosion ever recorded. Analysis of seismicity time-series before and after the Morakot typhoon reveals a systematic increase of shallow (i.e. 0-15 km of depth) earthquake frequency in the vicinity of the areas displaying a high spatial density of landslides. This step-like increase in frequency lasts for at least 2-3 years and does not follow an Omori-type aftershock sequence. Rather, it is associated to a step change of the Gutenberg-Richter b-value of the earthquake catalog. Both changes occurred in mountainous areas of southwest Taiwan, where typhoon Morakot caused extensive landsliding. These spatial and temporal correlations strongly suggest a causal relationship between the Morakot-triggered landslides and the increase of earthquake frequency and their associated b-value. We propose that the progressive removal of landslide materials from the steep mountain landscape by river sediment transport acts as an approximately constant increase of the stress rate with respect to pre-typhoon conditions, and that this in turn causes a step-wise increase in earthquake frequency. To test this hypothesis, we investigate the response of a rate-and-state fault to stress changes using a 2-D continuum elasto-dynamic model. Consistent with the results of Ader et al. (2013), our preliminary results show a step-like increase of earthquake frequency in response to a step-like decrease of the fault normal stress. We also investigate the sensitivity of the amplitude and time-scale of the earthquake frequency increase to the amplitude of the normal stress change and to rheological parameters. Our study offers new insights on the potential influence of extreme erosional events on the short-time scale dynamics of faults and earthquakes.
American Geophysical Union Fall Meeting 2016
San Francisco, United States

insu-01416987
https://hal-insu.archives-ouvertes.fr/insu-01416987

Éditeur(s) : HAL CCSD American Geophysical Union
Résumé : International audience
The impact of stratospheric ozone on the tropospheric general circulation of theSouthern Hemisphere (SH) is examined with a set of chemistry‐climate models participatingin the Stratospheric Processes and their Role in Climate (SPARC)/Chemistry‐ClimateModel Validation project phase 2 (CCMVal‐2). Model integrations of both the past andfuture climates reveal the crucial role of stratospheric ozone in driving SH circulationchange: stronger ozone depletion in late spring generally leads to greater polewarddisplacement and intensification of the tropospheric midlatitude jet, and greater expansion ofthe SH Hadley cell in the summer. These circulation changes are systematic as polewarddisplacement of the jet is typically accompanied by intensification of the jet and expansion ofthe Hadley cell. Overall results are compared with coupled models participating in theIntergovernmental Panel on Climate Change Fourth Assessment Report (IPCC AR4), andpossible mechanisms are discussed. While the tropospheric circulation response appearsquasi‐linearly related to stratospheric ozone changes, the quantitative response to a givenforcing varies considerably from one model to another. This scatter partly results fromdifferences in model climatology. It is shown that poleward intensification of the westerly jetis generally stronger in models whose climatological jet is biased toward lower latitudes.This result is discussed in the context of quasi‐geostrophic zonal mean dynamics.
ISSN: 2169-897X

hal-00337060
https://hal.archives-ouvertes.fr/hal-00337060
https://hal.archives-ouvertes.fr/hal-00337060/document
https://hal.archives-ouvertes.fr/hal-00337060/file/Bock_et_al-2008-Journalsur Geophysical_Research__Solid_Earth_%281978-2012%29.pdf
DOI : 10.1029/2008JD010327

Éditeur(s) : HAL CCSD
Résumé :  
The anatomy of tension wood and opposite wood was compared in 21 tropical rain forest trees from 21 species belonging to 18 families from French Guyana. Wood secimens were taken from the upper and lower sides of naturally titled trees. Measurement of the growth stress level ensured that the two samples were taken from wood tissues in a different mechanical state: highly tensile-stressed wood on the upper side, called tension wood and normally tensile-stressed wood on the lower side, called opposite wood. Quantitative parameters relating to fibres and vessels were measured on transverse sections of both tension and opposite wood to check if certain criteria can easily discriminate the two kinds of wood. We observed a decrease in the frequency of vessels in the tension wood in all the trees studied. Other criteria concerning shape and surface area of the vessels, fibre diameter or cell wall thickness did not reveal any general trend.At the ultrastructural level, we observed that the microfibril angle in the tension wood sample was lower than in opposite wood in all the trees except one (Licania membranacea).
IAWA Journal

hal-01032149
https://hal.archives-ouvertes.fr/hal-01032149

Éditeur(s) : HAL CCSD Oxford University Press (OUP)
Résumé : International audience
The interpretation of coseismic surface deformation measurements through inversion techniques is of major importance to understand the mechanical behaviour of a seismic fault. Dense geodetic data sets in the vicinity of the ruptured fault provide unique constraints on detailed fault geometry and slip distribution at depth, making them complementary to seismological data. Bam earthquake (Mw 6.6, 2003 December 26) induced surface deformation has been precisely mapped by Envisat ASAR interferometry and by subpixel correlation techniques applied to Spot-5 and ASAR amplitude images. These oblique and horizontal estimations of deformation have been completed with one levelling profile along the main road crossing the rupture from west to east. We process these data (separately and jointly) in a two-step inversion technique, within the elastic half-space theory framework. Our objective is to determine the dislocation model at depth that satisfies simultaneously all the geodetic constraints. Also, we estimate the relative contribution of each geodetic data set to this inversion process.We first use a stochastic direct approach called neighbourhood algorithm in order to estimate the average characteristics of the rupture, and their relative uncertainty. Constraining in this way the geometry of the ruptured fault, we then linearize the inverse problem and compute the slip distribution on the fault using a standard weighted least-square technique, assuming the solution is smooth to some degree. At each step, we discuss the optimal models, their stability as well as the relative influence of each data set on the derived models parameters. Our preferred model reveals a shallow dislocation on a quasi-vertical fault, slightly dipping towards east. The slip vector has a strike-slip component as high as 2 m, while the dip-slip component seems negligible.However, the estimation of the resolution matrices emphasizes the fact that the details of deep fault slip distribution remain out of the scope of this ill-conditioned inverse problem. Yet, our preferred model suggests a main dislocation limited at depth between 1 and 6 km. By contrast, the aftershocks observed in the months following the earthquake are located just beneath the estimated main shock.
ISSN: 0956-540X

hal-00314173
https://hal.archives-ouvertes.fr/hal-00314173
https://hal.archives-ouvertes.fr/hal-00314173/document
https://hal.archives-ouvertes.fr/hal-00314173/file/GJI-2007-169-849-briole.pdf
DOI : 10.1111/j.1365-246X.2007.03358.x

Éditeur(s) : HAL CCSD Elsevier
Résumé : International audience
This paper provides a synthesis of current data and interpretations on the crustal structure of the Pyrenean-Cantabrian orogenic belt, and presents new tectonic models for representative transects. The Pyrenean orogeny lasted from Santonian (~84 Ma) to early Miocene times (~20 Ma), and consisted of a spatial and temporal succession of oceanic crust/exhumed mantle subduction, rift inversion and continental collision processes at the Iberia-Eurasia plate boundary. A good coverage by active-source (vertical-incidence and wide-angle reflection) and passive-source (receiver functions) seismic studies, coupled with surface data have led to a reasonable knowledge of the present-day crustal architecture of the Pyrenean-Cantabrian belt, although questions remain. Seismic imaging reveals a persistent structure, from the central Pyrenees to the central Cantabrian Mountains, consisting of a wedge of Eurasian lithosphere indented into the thicker Iberian plate, whose lower crust is detached and plunges northwards into the mantle. For the Pyrenees, a new scheme of relationships between the southern upper crustal thrust sheets and the Axial Zone is here proposed. For the Cantabrian belt, the depth reached by the N-dipping Iberian crust and the structure of the margin are also revised.The common occurrence of lherzolite bodies in the northern Pyrenees and the seismic velocity and potential field record of the Bay of Biscay indicate that the precursor of the Pyrenees was a hyperextended and strongly segmented rift system, where narrow domains of exhumed mantle separated the thinned Iberian and Eurasian continental margins since the Albian-Cenomanian. The exhumed mantle in the Pyrenean rift was largely covered by a Mesozoic sedimentary lid that had locally glided along detachments in Triassic evaporites. Continental margin collision in the Pyrenees was preceded by subduction of the exhumed mantle, accompanied by the pop-up thrust expulsion of the off-scraped sedimentary lid above. To the west, oceanic subduction of the Bay of Biscay under the North Iberian margin is supported by an upper plate thrust wedge, gravity and magnetic anomalies, and 3D inclined sub-crustal reflections. However, discrepancies remain for the location of continent-ocean transitions in the Bay of Biscay and for the extent of oceanic subduction. The plate-kinematic evolution during the Mesozoic, which involves issues as the timing and total amount of opening, as well as the role of strike-slip drift, is also under debate, discrepancies arising from first-order interpretations of the adjacent oceanic magnetic anomaly record.
ISSN: 0040-1951

insu-01683989
https://hal-insu.archives-ouvertes.fr/insu-01683989
https://hal-insu.archives-ouvertes.fr/insu-01683989/document
https://hal-insu.archives-ouvertes.fr/insu-01683989/file/teixell-tectono-2018.pdf
DOI : 10.1016/j.tecto.2018.01.009

Éditeur(s) : HAL CCSD Elsevier
Résumé : International audience
The analysis of three cataclastic band sets from Provence (France) reveals that the band density, their conjugate angles, their ratio of shear displacement to compaction, and the amount of cataclasis within the bands differ and can be expressed as functions of tectonic setting and petrophysical properties. We identify (1) a dense and closely spaced network of shear-enhanced (reverse) compaction bands; (2) a regularly spaced less dense network of reverse compactional shear bands; and (3) a localized network of normal shear bands. The field data show that strain localization is favored in an extensional regime and is characterized by shear bands with a large shear to compaction ratio and a small conjugate band angle. In contrast, distributed strain is favored in a contractional regime and is characterized by compactional bands with a low ratio of shear to compaction and a large conjugate band angle. To explain the mechanical origin of this strain localization, we quantify the yield strength and the stress evolution in extensional and contractional regimes in a frictional porous granular material. We propose a model of strain localization in porous sands as a function of tectonic stresses, burial depth, material properties, strain hardening and fluid pressure. Our model suggests that stress reduction, inherent to extensional regime, favors strain localization as shear bands, whereas stress increase during contraction favors development of compactional bands.
ISSN: 0191-8141

hal-00856921
https://hal.archives-ouvertes.fr/hal-00856921
DOI : 10.1016/j.jsg.2012.11.011

Résumé : Le cadre juridique des investissements étrangers en Algérie a été marqué par de profonds changements depuis la promulgation du premier code des investissements, en 1963. Cette mutation n’a pas toujours été en phase avec les processus de développement économique national et le changement du contexte international. Le modèle socialiste de la gestion des affaires économiques qui a prédominé à partir du milieu des années 60 jusqu'au milieu des années 80 a progressivement cédé la place à un retrait graduel de l'Etat du champ économique. On assiste, depuis, à une émergence d’un secteur privé qui tente toujours de trouver sa place, à une vague de privatisations, à une libéralisation du commerce extérieur et à un appel aux capitaux étrangers pour financer le développement économique.Cette volonté manifeste d’accéder à une économie de marché est encore plus perceptible dans le dernier code des investissements de 2001. Ces efforts de mise en conformité du droit interne des investissements avec le droit international des investissements sont relayés par un discours politique offensif.Néanmoins, cet environnement juridique incitatif perd de son attrait lorsqu’on le confronte, soit aux textes promulgués en matière de réglementation du commerce extérieur et la réglementation des changes, soit à l’environnement administratif dans lequel est mis en œuvre l’ensemble de ces textes de lois. Cette confrontation est particulièrement instructive quant à l’insuffisance de cohérence de la politique étatique suivie en matière d’investissement étranger. Ceci révèle la balance d’intérêt des autorités algériennes entre l’augmentation de la contribution des investissements étrangers au potentiel économique et technologique national et la sauvegarde de la souveraineté nationale.Ainsi, le questionnement de la politique législative algérienne d’investissement permet de comprendre les fondements de la doctrine algérienne des investissements étrangers. Cette compréhension favorise l’émergence d’une stratégie qui concilie attractivité et préservation des intérêts publics.
The legal framework for foreign investment in Algeria was marked by profound changes since the enactment of the first investment code in 1963 This change has not always been in line with the process of national economic development and change in the international environment . The socialist model of economic management that prevailed from the mid 60s to mid 80s gradually gave way to a gradual withdrawal of the state from the economic field. There has since been an emergence of a private sector that is still trying to find its place, a wave of privatization, liberalization of foreign trade and to rely on foreign capital to finance the development will économique.Cette Manifest access to a market economy is even more noticeable in the latest investment Code of 2001 These efforts compliance of domestic investment law with the international investment law are supported by a political speech offensif.Néanmoins, the legal environment incentive loses its appeal when confronted or to enactments in regulation of foreign trade and foreign exchange regulations, or administrative environment that is implemented in all these laws. This comparison is particularly instructive about the lack of consistency in state policy with respect to foreign investment. This reveals the balance of interest between the Algerian authorities increasing the contribution of foreign investment to national economic and technological potential and safeguarding sovereignty nationale.Ainsi, questioning the Algerian legislative investment policy allows us to understand the foundations of the Algerian doctrine of foreign investment. This understanding promotes the emergence of a strategy that balances attractiveness and preservation of public interest.

http://www.theses.fr/2012AGUY0785

Éditeur(s) : HAL CCSD Elsevier
Résumé : International audience
Ecological signs of Earth's biosphere forewarn an alarming trajectory towards a global mass-extinction. Assessing species performance and susceptibilities to decline is essential to comprehend and reverse this trend. Yet it is challenging, given difficulties associated with quantifying individual and population processes that are variable across time, space, and life-stages. We describe a new approach to estimating and comparing species performances that combines empirical data, a novel theoretical consideration of population dynamics, and modern statistics. Our approach allows for a more realistic continuous representation of individual performances along development stages while taking into account non-linearity, and natural variability as captured by spatio-temporally replicated observations. We illustrate its application in a coral meta-assemblage composed of populations of the three major reef-building taxa Acropora, Pocillopora, Porites. Using a unique set of highly replicated observations of individual coral dynamics under various environmental conditions, we show how taxa differ in their investment in recruitment and size-specific aptitude for growth and survival, notably through different use of clonal shrinkage, fragmentation, fission, and fusion processes. Our results reveal contrasting life-history trade-offs among taxa which, along with differing patterns of density-dependent recruitment, modulate species responses to decline. These differences in coral life history traits reflect opposing life-strategies, imply regulation at differing life-stages, and explain divergence in species trajectories. Our findings indicate a high potential for resilience in Pocillopora and Porites populations, thanks respectively to a sustained recruitment that promotes demographic elasticity through replacement of individuals, and a steady resistance to mortality which confers persistence through lingering of individuals. Resilience in Acropora appears more arbitrary, given high susceptibility to perturbations and dependency of recruitment on presence of established local populations. We identify management actions that can complement Acropora's life history and benefit recovery of its populations following mortality events. Our regression-modelling approach to quantifying and comparing species performances in different population processes is applicable to all taxa, as illustrated even those with complex clonal life histories, and can be implemented at wide spatio-temporal and taxonomic coverage. It can promote more accurate representation of species dynamics in both descriptive and predictive modelling approaches. The semi-parametric contrast curve method we develop facilitates comparing response variables along continuous explicative metrics while accounting for multiple sources of complexity in empirical data. It should widely benefit investigations in ecology and quantitative science.
ISSN: 1476-945X

hal-01212337
https://hal-univ-perp.archives-ouvertes.fr/hal-01212337
DOI : 10.1016/j.ecocom.2015.07.003

Éditeur(s) : HAL CCSD American Geophysical Union (AGU)
Résumé : International audience
The Atlas Mountains are characterized by high elevations and Quaternary volcanism. Long period magnetotelluric data acquired along a NNW-SSE transect reveal the presence of a conductive anomalous mantle below the High Atlas. Data dimensionality analyses show a preferent N80 degrees E strike of the deep resistivity structure in agreement with the induction vector alignment at long periods. Accordingly, a 2D inversion of the data set was carried out. Large resistive bodies at the crustal basement most likely correspond to batholiths emplaced in more conductive metapelites. They are covered by outcropping conductive sedimentary detritic and carbonate rocks. Lithospheric thinning producing anomalous mantle and basin development in the Atlas probably started during Triassic-Jurassic rifting. Inversion tectonics since the Oligocene produced low shortening on previous lithospheric weak zones, with thrusting of the Atlas above the stable African plate. Melting at the top of the anomalous mantle is connected with Quaternary basaltic volcanism in the Middle Atlas.
ISSN: 0278-7407

hal-00671017
https://hal.archives-ouvertes.fr/hal-00671017
DOI : 10.1029/2010TC002859

Éditeur(s) : HAL CCSD American Geophysical Union
Résumé : International audience
Torsion experiments (inline image =3 × 10−4 s−1) were performed to investigate the large strain (γmax = 15) rheology on quartz-muscovite aggregate as analogue to pelitic rocks undergoing melting and crystallization during deformation at 300 MPa confining pressure and 750°C temperature. Microstructures reveal four distinct but gradational stages of crystal-melt interactions during deformation--(a) solid state deformation, (b) initiation and domination of partial melting, (c) simultaneous partial melting and crystallization, and (d) domination of crystallization. The microstructural stages are linked to the rheology of the deforming samples. Partial melting starts at relatively low finite shear strains (γ = 1-3) showing approximately 60% strain softening. At γ = 4-10 the partially molten bulk material shows a steady state flow at low stress. Further crystallization of new crystals at the expense of melt between γ = 10 and 15 causes weak strain hardening until the material fails by developing brittle fractures. The stress exponent (n), calculated at γ = 1, 5, and 10, increases from ∼ 3 to ∼ 43, indicating a transition from power to power law breakdown or exponential flow of the bulk system. Hydrostatic experiments for equivalent times and conditions of the torsion experiments were also conducted to evaluate the reaction kinetics and microstructures under static conditions. The new experimental data establish that partially molten rock does not flow according to a constant strain rate-dependent power law (steady state) rheology. The rheological transition from strain rate sensitive to strain rate insensitive flow is interpreted as a function of melt-crystal ratio, their mutual interactions, and the evolution of microstructures in the partially molten rock.
ISSN: 0148-0227

hal-01053709
https://hal.archives-ouvertes.fr/hal-01053709
DOI : 10.1002/2013JB010777

Éditeur(s) : HAL CCSD Springer Verlag
Résumé : International audience
Standard microbial evolutionary ontology is organized according to a nested hierarchy of entities at various levels of biological organization. It typically detects and defines these entities in relation to the most stable aspects of evolutionary processes, by identifying lineages evolving by a process of vertical inheritance from an ancestral entity. However, recent advances in microbiology indicate that such an ontology has important limitations. The various dynamics detected within microbiological systems reveal that a focus on the most stable entities (or features of entities) over time inevitably underestimates the extent and nature of microbial diversity. These dynamics are not the outcome of the process of vertical descent alone. Other processes, often involving causal interactions between entities from distinct levels of biological organisation, or operating at different time scales, are responsible not only for the destabilisation of pre-existing entities, but also for the emergence and stabilisation of novel entities in the microbial world. In this article we consider microbial entities as more or less stabilised functional wholes, and sketch a network-based ontology that can represent a diverse set of processes including, for example, as well as phylogenetic relations, interactions that stabilise or destabilise the interacting entities, spatial relations, ecological connections, and genetic exchanges. We use this pluralistic framework for evaluating (i) the existing ontological assumptions in evolution (e.g. whether currently recognized entities are adequate for understanding the causes of change and stabilisation in the microbial world), and (ii) for identifying hidden ontological kinds, essentially invisible from within a more limited perspective. We propose to recognize additional classes of entities that provide new insights into the structure of the microbial world, namely ``processually equivalent'' entities, ``processually versatile'' entities, and ``stabilized'' entities.
ISSN: 0169-3867

hal-01544796
https://hal.archives-ouvertes.fr/hal-01544796
DOI : 10.1007/s10539-012-9350-2

Éditeur(s) : HAL CCSD Elsevier
Résumé : The variability of fluvial activity in the Northwestern Mediterranean coastal lowlands and its relationship with modes of climate change were analysed from the late 9th to the 18th centuries CE. Geochemical analyses were undertaken from a lagoonal sequence and surrounding sediments in order to track the fluvial inputs into the lagoon. An index based on the K/S and Rb/S ratios was used to evidence the main periods of fluvial activity. This index reveals that the Medieval Climate Anomaly (MCA) was a drier period characterized by a lower fluvial activity, while the Little Ice Age (LIA) was a wetter period with an increase of the river dynamics. Three periods of higher than average fluvial activity were evidenced at the end of the first millennium CE (ca. 900–950 cal yr CE), in the first half of the second millennium CE (ca. 1150–1550 cal yr CE), and during the 1600s–1700s CE (ca. 1650–1800 cal yr CE). The comparison of these fluvial periods with other records of riverine or lacustrine floods in Spain, Italy, and South of France seems to indicate a general increase in fluvial and flood patterns in the Northwestern Mediterranean in response to the climate change from the MCA to the LIA, although some episodes of flooding are not found in all records. Besides, the phases of higher than average fluvial dynamics are in good agreement with the North Atlantic cold events evidenced from records of ice-rafted debris. The evolution of fluvial activity in the Northwestern Mediterranean coastlands during the last millennium could have been driven by atmospheric and oceanic circulation patterns.Keywords
ISSN: 0921-8181

hal-01571958
https://hal.archives-ouvertes.fr/hal-01571958
DOI : 10.1016/j.gloplacha.2017.03.008

Éditeur(s) : HAL CCSD Elsevier
Résumé : International audience
In order to estimate the seismic anisotropy of subducted crust, polycrystalline samples of KA1Si(3)O(8) K-lingunite (25% of the total subducted transformed sediments), have been synthesized and deformed under the temperature and pressure conditions of the subducted slabs. Transmission electron microscopy (TEM) characterizations of the recovered samples reveal that the microstructures are clearly dominated by [001] glide involving screw dislocations. For this reason, only {100} could be identified as glide planes, the question of [001] slip on {110} remains open. Few 1/2 < 111 > dislocations were observed gliding on {110} planes, which implies that 1/2 < 111 > {110} is a harder slip system than those involving [001] slip. The occurrence of sub-grain boundaries suggests that diffusion and climb might be active under these conditions.;To assess the texture of polycrystalline K-lingunite, the crystal preferred orientations (CP05) were calculated using visco-plastic self-consistent (VPSC) polycrystalline plasticity model in simple shear using the slip systems identified by TEM. Finally, the seismic properties of K-lingunite aggregates were calculated from the CPO and single crystal elasticity tensor. K-lingunite is predicted to have a high seismic anisotropy, which could combine constructively with one of the stishovite (same proportion as K-lingunite at the transition zone depth ranges).
ISSN: 0031-9201

hal-00534050
https://hal.archives-ouvertes.fr/hal-00534050
DOI : 10.1016/j.pepi.2010.06.013

Résumé : Nous avons étudié les phénomènes de polyploïdisation chez l'igname Dioscorea alata qui comprend trois cytotypes diploïde (2n=40), triploïde (2n=60) et tétraploïde (2n=80) en vue d'optimiser les stratégies de production d'hybrides polyploïdes. Chez ce complexe polyploïde l'augmentation de la ploïdie est corrélée avec une augmentation de la vigueur et des rendements plus élevés et plus stables. Dans un premier temps nous avons démontré l'autotétraploïdie des variétés tétraploïdes à l'aide de trois approches différentes: des analyses d'hérédité de marqueurs microsatellites, l'observation de phénomènes de double réduction et l'étude des méioses des cellules mères de pollen. Nous avons ensuite déterminé quels sont les mécanismes les plus probables à l'origine des polyploïdes naturels via l'étude de la transmission de l'hétérozygotie parentale à l'aide de microsatellites et par l'étude des incompatibilités au niveau de l'albumen lors de différents croisements intracytotypes et intercytotypes en utilisant la cytométrie en flux. Les résultats obtenus montrent que les polyploïdes de D. alata seraient apparus via la formation de gamètes non réduits de clones diploïdes. Le pool triploïde se serait édifié et diversifié uniquement par voie femelle, du fait de la non viabilité des croisements intercytotypes et de la stérilité des femelles et mâles triploïdes. Le pool tétraploïde serait apparu par union de deux gamètes non réduits de clones diploïdes (polyploidisation sexuelle bilatérale). Par la suite ce pool aurait été diversifié via des croisements intercy
We studied polyploidisation phenomena in the yam Dioscorea alata that includes three cytotypes -diploid (2n=40), triploid (2n=60) and tetraploid (2n=80) -in order to optimise polyploid hybrid production strategies. In this complex polyploid, the increase in ploidy is correlated with an increase in vigour and higher and more stable yields. We first showed the autotetraploidy of tetraploid varieties using three different approaches: heredity analyses of microsatellite markers, the observation of double reduction phenomena, and the study of meiosis of pollen mother cells. We then determined the mechanisms most likely to be at the origin of natural polyploids through the study of the transmission of parental heterozygoty using microsatellites and the study of incompatibilities at the endosperm level at the time of different intracytotypic and intercytotypic crosses using flow cytometry. The results obtained reveal that the polyploids of D. alata probably appeared through the formation of non-reduced gametes of diploid clones. The triploid pool would then have been constituted and diversified through the female pathway as a result of the non-viability of intercytotypic crosses and the sterility of female and male triploids. The tetraploid pool would have appeared as a result of the union of two non-reduced gametes of diploid clones (bilateral sexual polyploidisation). This pool would then have diversified through intercytotypic crosses with the formation of 2n gamètes through both the female and the male pathway, as well as by intracytotypic crosses within the 4X pool.

http://www.theses.fr/2012AGUY0519/document