Éditeur(s) : HAL CCSD Elsevier
Résumé : International audience
The Massif du Sud is a large ophiolitic complex that crops out in the southern region of New Caledonia (SW Pacific). It is dominated by harzburgite tectonite that locally shows a transitional gradation to massive dunite up section. Clinopyroxene, orthopyroxene and plagioclase progressively appear in dunite up to the transition to layered wehrlite and orthopyroxene-gabbro. The dunite-wehrlite and wehrlite-gabbro contacts are parallel and the latter defines the paleo-Moho.;Highly depleted modal, mineral and bulk rock compositions indicate that harzburgites are residues after high degrees (20-30%) of partial melting mainly in the spinet-stability field. Their relative enrichment in HFSE, LREE and MREE is due to re-equilibration of melting residues with percolating melts. Dunite formed in the Moho transition zone by reaction between residual mantle harzburgite and olivine-saturated melts that led to pyroxene dissolution and olivine precipitation. Rare clinopyroxene and plagioclase crystallized in interstitial melt pores of dunite from primitive, low-TiO2, ultra-depleted liquids with a geochemical signature transitional between those of island arc tholeiites and boninites.;Ascending batches of relatively high-SiO2, ultra-depleted melts migrated through the Moho transition zone and generated wehrlite by olivine dissolution and crystallization of clinopyroxene, orthopyroxene and plagioclase in variable amounts. These liquids were more evolved and were produced by higher degrees of melting or from a more depleted source compared with melts that locally crystallized clinopyroxene in dunite. Ultra-depleted magmas, non-cogenetic with those that formed the Moho transition zone, ascended to the lower crust and generated gabbroic cumulates with subduction-related affinity. Thus, the ultramafic and mafic rocks in the Moho transition zone and lower crust of the Massif du Sud ophiolite are not products of fractional crystallization from a single magma-type but are the result of migration and accumulation of different melts in a multi-stage evolution. The record of high partial melting in the mantle section, and migration and accumulation of ultra-depleted subduction-related melts in the Moho transition zone and lower crust support that the Massif du Sud ophiolite is a portion of forearc lithosphere generated in an extensional regime during the early phases of the subduction zone evolution. Our results show the existence of different types of ultra-depleted melt compositions arriving at the Moho transition zone and lower crust of an infant intraoceanic paleo-arc. Ultra-depleted melts may thus be a significant component of the melt budget generated in oceanic spreading forearcs prior to aggregation and mixing of a large range of melt compositions in the crust.
ISSN: 0009-2541

hal-00429333
https://hal.archives-ouvertes.fr/hal-00429333
DOI : 10.1016/j.chemgeo.2009.06.004

Éditeur(s) : HAL CCSD EDP Sciences
Résumé : International audience
In this work, we point out and apply some ideas for the numerical simulation by multi- level schemes of several wave equations including dispersive (Korteweg de Vries) and dissipative ones (Kuramoto-Sivashinski). We start from the approach developed for dissipative equations decomposing the solution into large (Y) and small (Z) scales, and we show that it is necessary to have a regularization in time to allow stable numerical treatment of Y and Z which are here generated using hierarchical processes such as Wavelets, Incremental Unknowns or Fourier.
ISSN: 1270-900X

hal-00768332
https://hal.archives-ouvertes.fr/hal-00768332
https://hal.archives-ouvertes.fr/hal-00768332/document
https://hal.archives-ouvertes.fr/hal-00768332/file/esaim_proc_2009_v27.pdf

Éditeur(s) : HAL CCSD Elsevier
Résumé : International audience
Reef positions record the interaction between eustasy and tectonics, and have been used worldwide to characterize vertical deformations of upper-plates at different time-scales and constrain the seismic behavior of megathrusts. Along the Lesser Antilles volcanic arc, high-resolution marine geophysical data were collected on the 2–20 km wide eastern Martinique reef platform to reconstruct its stratigraphic and morphologic history, and understand the influence of local normal faulting, volcanism and plate-scale subduction processes on Holocene and Late Pleistocene reef development. The subsiding Martinique platform's stratigraphy is composed of multiple superimposed sea-level highstand deposits separated by subaerial exposure surfaces of sea-level low stands. The carbonate platform consists of two laterally-extensive carbonate units (unit U2 overlying unit U3) that extend to the platform edge to a depth of − 95 m MSL (mean sea level), and form two morphologic terraces, M2 and M3 respectively. The landward portion of unit U2 is partially overlain between 0 and − 60 m MSL by the living reef tract U1. The current reef is composed of a landward fringing reef, a lagoon and a seaward barrier reef, the latter forming a double-bank barrier around the Caravelle Peninsula. In near-shore multi-channel seismic profiles, a distinct reflector at ~− 35 m MSL, probably a subaerial exposure surface E1, separates the reef sequence formed during the last transgression from a Pleistocene fossil reef tract forming unit U2. Offshore of Mount Pelée volcano (Late Pleistocene), the Holocene reef did not develop above unit U2, whose upper surface is incised by channels and apparent sinkholes. During the Holocene transgression, the possibility of excessive turbidity due to volcanic activity may have inhibited reef development in this area. The un-dated unit U2 probably developed 120–130 ka ago during the last interglacial (MIS 5.5) + 6 m MSL highstand as thick, extensive reefs deposited all along the Lesser Antilles arc. Due to subsidence, MIS 5.5 reefs are not represented by onshore facies, except along the southern Sainte Anne Peninsula where normal faulting and uplift balances island-scale subsidence. Based on unit U2's present elevation and assuming an MIS 5.5 age and + 6 m MSL sea level, Martinique has subsided at maximum 0.3 m/ky, likely due to subduction processes that question the coupling state of the megathrust.
ISSN: 0025-3227

hal-01307036
https://hal.archives-ouvertes.fr/hal-01307036
DOI : 10.1016/j.margeo.2015.08.001

Éditeur(s) : HAL CCSD Elsevier
Résumé : International audience
Wallace (1951) and Bott (1959) were the first to introduce the idea that the slip on each fault surface has the same direction and sense as the maximum shear stress resolved on that surface. However, this simplified hypothesis is questionable since fault mechanical interactions may induce slip reorientations. Earlier numerical geomechanical models confirmed that the slickenlines (slip vectors) are not necessarily parallel to the maximum resolved shear stress but are consistent with local stress perturbations. This leads us to ask as to what extent the Wallace and Bott simplifications are reliable as a basis hypothesis for stress inversion from fault slip data. Here, a geomechanical multi-parametric study using a 3D boundary element method, covering (i) fault geometries such as intersected faults or corrugated fault surfaces, (ii) the full range of Andersonian state of stress, (iii) fault friction, (iv) fault fluid pressure, (v) half space effect and (vi), rock properties, is performed in order to understand the effect of each parameter on the misfit angle between geomechanical slip vectors and the resolved shear stresses. It is shown that significant misfit angles can be found under specific configurations invalidating the Wallace and Bott assumptions, even though fault friction tends to minimize the misfit. We therefore conclude that in such cases, stress inversions based on fault slip data should be interpreted with care.
ISSN: 0040-1951

hal-01218418
https://hal.archives-ouvertes.fr/hal-01218418
DOI : 10.1016/j.tecto.2015.06.030

Résumé : La nécessité d'une approche intégrée des pêches est actuellement largement affirmée, notamment par la FAO, en particulier dans le contexte des pêcheries artisanales tropicales. Ces pêcheries à petites échelles, souvent multi-spécifiques sont très importantes en termes d'emploi et de production, y compris pour la sécurité alimentaire. Néanmoins ces pêcheries et la biodiversité exploitées sont souvent marquées, d'une part, par le manque de données rendant difficile leur gestion dans le cadre d'une approche écosystémique, d'autre part, par la complexité des systèmes écologiques et économiques sous-jacents incluant interactions trophiques et techniques.Cette thèse contribue à la mise au point d'outils bioéconomiques adaptés aux petites pêcheries tropicales dans la perspective d'une gestion durable des pêches fondée sur l'approche écosystémique. Dans cette perspective, la pêcherie côtière en Guyane Française constitue un cas d'étude particulièrement fécond. La thèse propose ainsi des évaluations multi-critères, des modèles bioéconomiques complexes et des scénarios de gestion viable pour cette pêcherie.Dans un premier temps l'analyse statistique multi-variée suggère un statut global satisfaisant de la pêcherie en termes de durabilité. Cependant des différences de performance sont notées au sein de la pêcherie, notamment au niveau des zones frontalières. Aussi des améliorations dans le mode de gestion sont proposées. Ensuite, à travers la modélisation bioéconomique, les projections des différents scénarios de pêche montrent qu'à long terme le niveau d'exploitation actuel peut ne pas être en adéquation avec la future augmentation de la demande locale et qu'une perte de biodiversité peut avoir lieu. Un scénario dit de « co-viabilité » conciliant des objectifs à la fois écologique, économique et social, avec une probabilité de réalisation satisfaisante, est mis en exergue. Enfin, la comparaison des comportements optimaux en situation coopératif et non coopératif, montre dans quelle mesure la viabilité est favorisée quand les acteurs coopèrent. Au-delà du diagnostic apporté pour le cas d'étude, la méthode utilisée s'avère prometteuse dans le contexte des petites pêcheries tropicales, tandis que l'approche de « co-viabilité » permet de trouver les modalités d'exploitation dans le cadre de compromis entre durabilité écologique et socio-économique, ce que les principes de gestion halieutiques traditionnels ne permettent généralement pas.
The need for an integrated approach to fisheries is now widely affirmed, including the FAO, in particular in the context of tropical artisanal fisheries. These small-scale fisheries, often multispecies are very important in terms of production, employment and food security. However these fisheries and the exploited resources are often characterized by a lack of data making difficult their management within an ecosystem approach, and by the ecological and economic system complexities including trophic and technological interactions.This thesis aims to contribute to the development of bioeconomic tools for small tropical fisheries in a sustainable management perspective based on the ecosystem approach. In this perspective, the French Guiana coastal fishery constitutes an interesting case study. The thesis proposes multi-criteria evaluations, complex bioeconomic models and viable management scenarios for this fishery. At first, multivariate statistical analysis suggests a satisfactory overall status of the fishery in terms of sustainability. However, some performance differences are noted within the fishery, particularly at border areas. Also, management improvements are proposed. Then, through bioeconomic modeling, projections of different fishing scenarios show that, in the long term, the current exploitation level may not be consistent with the future increase of local demand and a loss of biodiversity may occur. A scenario called "co-viability" reconciling ecological, economic and social objectives, with a high probability of achievement is exhibited. Finally, the comparison of optimal behavior in cooperative and non-cooperative conditions, confirms that harvest levels are greater when actors do not cooperate. Furthermore, it is shown that the state of the ecosystem depends on the fishing strategies and the type of interaction between species. Beyond the diagnosis made for the case study, the method is promising in the context of small tropical fisheries, while the co-viability approach allows finding the exploitation conditions under which ecological and socio-economic sustainability are meet, what the conventional fisheries management generally do not allow.

http://www.theses.fr/2013AGUY0620/document

Éditeur(s) : Elsevier Sci Ltd
Résumé : A multi-scale global hindcast of ocean waves is presented that covers the years 1994-2012, based on recently published parameterizations for wind sea and swell dissipation [Ardhuin, F., Rogers, E., Babanin, A., Filipot, J.-F., Magne, R., Roland, A., van der Westhuysen, A., Queffeulou, P., Lefevre, J.-M., Aouf, L., Collard, F., 2010. Semi-empirical dissipation source functions for wind-wave models: Part I. Definition, calibration and validation. J. Phys. Oceanogr. 40 (9), 1917-1941]. Results from this hindcast include traditional wave parameters, like the significant wave height and mean periods, and we particularly consider the accuracy of the results for phenomenal sea states, with significant heights above 14 m. Using unbiased winds, there is no evidence of a bias in wave heights even for this very high range. Various spectral moments were also validated, including the surface Stokes drift and mean square slopes that are relevant for wave-current interactions modelling and remote sensing, and also spectra of seismic noise sources. The estimation of these parameters is made more accurate by the new wave growth and dissipation parameterizations. Associated air-sea fluxes of momentum and energy are significantly different from what is obtained with the WAM-Cycle 4 parameterization, with a roughness that is practically a function of wind speed only. That particular output of the model does not appear very realistic and will require future adjustments of the generation and dissipation parameterizations.
Ocean Modelling (1463-5003) (Elsevier Sci Ltd), 2013-10 , Vol. 70 , P. 174-188

Droits : 2012 Elsevier Ltd. All rights reserved.
http://archimer.ifremer.fr/doc/00155/26582/25477.pdf
DOI:10.1016/j.ocemod.2012.12.001
http://archimer.ifremer.fr/doc/00155/26582/

Résumé : Les pathologies inflammatoires démyélinisantes primitives du système nerveux central (SNC) sont dominées par deux entités : la sclérose en plaques (SEP) et la neuromyélite optique (NMO). Ces deux pathologies se différencient par : 1. leur phénotype, la NMO se caractérisant par une myélite extensive dans le plan transversal et longitudinal et peu de lésions encéphaliques initiales ; 2. leur sévérité, le pronostic de la NMO étant moins bon que celui de la SEP ; 3. leur mécanisme immunologique, puisque l’immunité humorale domine nettement dans la NMO, cette pathologie répondant peu ou pas aux traitementshabituellement prescrits dans la SEP. Un anticorps sérique (IgG-NMO) a été retrouvé chez les patients atteints de NMO avec une sensibilité de 50-70% et une spécificité de 90-100%. Cet anticorps a pour cible le principal canal hydrique du SNC, l’aquaporine-4 (AQP4), qui se situe principalement sur les pieds astrocytaires jalonnant la barrière hémato-encéphalique. La SEP est relativement familière des neurologues car décrite sur le plan épidémiologique et dans son histoire naturelle depuis plus d’un siècle à travers de larges séries réalisées parfois en population générale. Son approche thérapeutique progresse suite à des essais contrôlés qui s’accumulent depuis maintenant près de 20 ans. Toutefois, la majorité des travaux ont été effectués chez les populations Caucasiennes en raison de sa plus grande fréquence dans ces populations. A l’inverse, la NMO garde encore un relatif mystère à cause de sa relative rareté, l’établissement très récent de critères diagnostiques et une physiopathologie très récemment éclaircie par la découverte de l’IgG-NMO qui a indiscutablement suscité un regain d’intérêt de cette affection. Ainsi, son incidence est quasiment inconnue de même que sa susceptibilité génétique. La progression du handicap de la NMO qu’il soit visuel ou moteur à l’évidence sévère, ne repose toujours pas sur des chiffres précis, la description des lésions neuroradiologiques reste encore très embryonnaire. Enfin, même s’il existe un consensus pour traiter énergiquement les poussées de NMO et les espacer, il n’existe quasiment aucune étude prospective.
Introduction: Aim of the study is to describe épidemiology, phenotype and treatment of demyelinating inflammatory diseases of the central nervous system (CNS) namely multiple sclerosis (MS) and neuromyelitis optica (NMO) in French West Indies.Epidemiology: MS emerged in the French West Indian (FWI) population with an incidence of 1.27/100 000 (CI 95%: 1.16-1.38) for the period 1992-2007 while incidence of NMO remained stable at 0.19/100,000 (CI 95%: 0.15–0.23). Both migration in Metropolitean France (MF) and decrease of sun exposure were risk factors for MS acquisition in FWI. Phenotype and treatment of NMO: NMO is a very rapidly disabling disease with median times to reach DSS 3, 6 and 8 of one year, 8 years and 22 years respectively. However, treatment by plasma exchanges were shown to reduce sequellae of NMO attacks ; mitoxantrone and rituximab reduced significantly annualized relapse rate (ARR) from 1.7 to 0.29 (p<10-4) and from 1,34 to 0,56 (p=0.0005) respectively, improving dramatically natural course of NMO.Phenotype and treatment of MS: Absence of Migration in MF in MS patients was found to be the strongest independent predictive factor of disability progression to DSS 6 (hazard ratio, 2.59; p=0.0002); beside short time to DSS3 (p=0.005), later age at onset (p=0.005), and multi-symptomatic onset (p=0.03). Treatment of MS by interferon beta-1 had a moderate impact in MS while natalizimab proved to be highly effective by reducing drastically ARR and inflammatory parameters on magnetic resonance imaging. Conclusion: Spectrum of demyelinating inflammatory disorders of CNS has reversed in French West Indies with overrepresentation of MS. Agressive therapy is needed to improve prognosis of NMO. Rapidly increasing of its incidence makes MS a new public health problem in French West Indies.

http://www.theses.fr/2012AGUY0511/document

Éditeur(s) : HAL CCSD Springer Verlag
Résumé : Some major challenges for geophysicists and structural geologists using three-dimensional boundary element method codes (3D-BEM) are: (1) reducing the amount of memory required to solve large and dense systems and (2) incorporation of inequality constraints such as traction inequality constraints (TIC) and displacement inequality constraints (DIC). The latter serves two purposes. First, for example, inequality constraints can be used to simulate frictional slip (using TIC). Second, these constraints can prevent element interpenetration while allowing opening mode (using DIC). We have developed a method that simultaneously incorporates both types of functionality of the inequality constraints. We show that the use of an appropriate iterative solver not only avoids the allocation of significant memory for solving the system (allowing very large model computation and simplifying parallelization on multi-core processors), but also admits interesting features such as natural incorporation of TICs and DICs. Compared to other techniques of contact management (e.g., Lagrange multipliers, penalty method, or complementarity problem), this new simple methodology, which does not use any incremental trial-and-error procedures, brings more flexibility, while making the system more stable and less subject to round-off errors without any computational overhead. We provide validations and comparisons of the inequality constraints implementation using 2D analytical and numerical solutions.
ISSN: 1420-0597

hal-00561035
https://hal.archives-ouvertes.fr/hal-00561035
DOI : 10.1007/s10596-009-9170-x

Éditeur(s) : HAL CCSD Elsevier
Résumé : International audience
Carbonation of peridotite is a widespread process in nature, with emerging societal and environmental implications through geological storage of CO2. We studied the micro- and nano-scale structures associated with the carbonation of peridotite pervaded by a CO2-rich water and the induced changes of its hydrodynamic properties. Focused ion beam (FIB) nanotomography was coupled with scanning electron microscopy (SEM) and transmission electron microscopy (TEM) observations. An experimental sample of dunite core percolated by a CO2-rich fluid (pCO2 = 11 MPa) at 160 °C was compared with a naturally carbonated sample from the Atlantic Ridge (Exp. IODP 304). Results of the 8-days percolation experiment show continuous alteration of olivine into magnesite locally associated with a poorly crystalline serpentine-type mineral referred to as proto-serpentine. The experiment is also characterized by an overall decrease in rock porosity (from 7.7 to 6.4%) and permeability (from 8.5 × 10− 16 to 1.7 × 10− 16 m2). The primary inter-granular porosity (1 to 10 μm) is progressively filled with magnesite, leading to the overall decrease of permeability. At the same time, a secondary small-scale porosity (10 nm to 1 μm) is created at the tip of the dissolution features along the (010) cleavage planes of olivine grains. Carbonate crystals grow within these etch-pit structures behind the dissolution front. Dissolution and precipitation occur at a relative rate that maintains the secondary porosity over the timescale of the experiment (i.e. carbonate growth has to be the slowest one). This process is interpreted as an interface coupled dissolution-precipitation mechanism of olivine carbonation. Proto-serpentine is only formed in highly reduced flow areas, such as small olivine etch-pits (10 to 500 nm wide), which are more isolated from CO2 input. SEM-FIB analyses show the 3D preferential orientation of the dissolution-precipitation planes within a given olivine grain. It suggests the formation of lateral connections between the etch-pits (infra-micrometric “channels”) providing access to reactants and removing reaction products at the reaction sites. The experimental results show that olivine dissolution and magnesite precipitation converge to a quasi-steady state resulting in a constant carbonation rate controlled by reaction kinetics and not by mass transfer for this flow regime. The system self-regulates, independently of the porosity change at least during this incipient stage of olivine carbonation. Similar textures are observed in natural samples collected at 170 m-depth in the vicinity of veinlets driving late fluids down into the cooling oceanic lithosphere. This suggests that similar mechanisms are active in nature and can play an important role in the late, low temperature alteration stages of fractured peridotite. These results also highlight the critical role of nm-scale spatial arrangement of dissolution and precipitation features on mass transfers and on the sustainability of olivine carbonation through coupled dissolution-precipitation processes.
ISSN: 0009-2541

hal-01684733
https://hal.archives-ouvertes.fr/hal-01684733
DOI : 10.1016/j.chemgeo.2017.11.013

Éditeur(s) : HAL CCSD Elsevier
Résumé : International audience
Four decades after their introduction, the validity of fault slip inversion methods based on Wallace (1951) and Bott (1959) hypothesis, which states that the slip on each fault surface has the same direction and sense as the maximum resolved shear stress, is still a subject of debate. According to some authors, this hypothesis is questionable since fault mechanical interactions induce slip reorientations as confirmed by geomechanical models. This leads us to ask as to what extent the Wallace-Bott simplifications are reliable as a basis hypothesis for stress inversion from fault slip data. In this paper, we compare two inversion methods; the first is based on the Wallace-Bott hypothesis, and the second relies on geomechanics and mechanical effects on fault heterogeneous slip distribution. In that context, a multi-parametric stress inversion study covering (i) the friction coefficients (μ), (ii) the full range of Andersonian state of stress and (iii) slip data sampling along the faults is performed. For each tested parameter, the results of the mechanical stress inversion and the Wallace-Bott (WB) based stress inversion for slip are compared in order to understand their respective effects. The predicted discrepancy between the solutions of both stress inversion methods (based on WB and mechanics) will then be used to explain the stress inversions results for the chimney Rock case study. It is shown that a high solution discrepancy is not always correlated with the misfit angle (ω ) and can be found under specific configurations (View the MathML sourceR−, θ, μ, geometry) invalidating the WB solutions. We conclude that in most cases the mechanical stress inversion and the WB based stress inversion are both valid and complementary depending on the fault friction. Some exceptions (i.e. low fault friction, simple fault geometry and pure regimes) that may lead to wrong WB based stress inversion solutions are highlighted.
ISSN: 0040-1951

hal-01467165
https://hal.archives-ouvertes.fr/hal-01467165
DOI : 10.1016/j.tecto.2016.11.039

Éditeur(s) : HAL CCSD Coastal Education and Research Foundation
Résumé : Nearshore bars are generally present on sandy coasts and play a significant role in wave breaking and sediment exchange between beach and shoreface. The study area is located on the Languedoc-Roussillon coast in the Gulf of Lions (southern part of the French Mediterranean coast). This microtidal environment is classified as a wave-dominated coast. The sand coast displays different bar systems (single or multiple; straight or crescentic). The alongshore variability of bar morphology in the nearshore zone was investigated in August 2009 during one week, using a topo-bathymetric LIDAR data set (300 km(2)). LIDAR makes it possible to cover a large area in a very short time with a high resolution. The topo-bathymetric LIDAR offers the added benefit of morphological information about the beach-sea transition in very shallow water (below I in), data often difficult if not impossible to obtain with traditional techniques. The purpose of this paper is 1) to depict the distribution of bars and their characteristics along the 180 km coast of the Gulf of Lions; 2) to update the classifications of sand bars typologies in Languedoc-Roussillon; 3) to pay particular attention to the description of the intricate inner system. Globally, cross-shore distribution of subtidal morphologies is characterized by both an outer and an inner system. The outer system is crescent-shaped in the south and straight in the northern part with a distinctive regional delimitation. The inner system comprises an inner bar and, near the coast, a complex fluctuating bar called Low Bar Beach (LBB). Several intermediate states were identified and added to the new classification proposed in this paper. These results clearly emphasize the importance of LIDAR technology for a better understanding of bar behaviour and single and/or multi-bar beach organisation.
ISSN: 0749-0208

hal-00783187
https://hal.archives-ouvertes.fr/hal-00783187

Résumé : Les mutualismes, interactions interspécifiques où chaque partenaire retire un bénéfice net de leur association, sont centraux dans l’origine et l’organisation de la biodiversité. Bien que globalement bénéfiques pour chacun des partenaires, ces interactions n’enlèvent rien à l’égoïsme inhérent de chaque espèce pour sa survie et sa reproduction, générant des conflits d’intérêts entre les espèces. Ainsi, comprendre les processus écologiques et évolutifs qui maintiennent le caractère mutualiste d’interactions entre plusieurs espèces est primordial dans la compréhension du maintien de la biodiversité. Néanmoins, le corpus scientifique s’est jusqu'à présent surtout concentré sur des paires d’espèces en interaction. Or, ces avancées scientifiques restent partielles car la plupart de ces interactions s’englobent dans un contexte communautaire. C’est dans ce cadre conceptuel que se place mon travail de thèse. Alors que la diversité structurelle des mutualismes de protection entre plantes et fourmis en ont fait un modèle d’étude clé dans la compréhension des mutualismes, j’ai concentré mes recherches sur l’intégration d’un troisième partenaire fongique pour comprendre ses conséquences sur les résultantes écologiques et évolutives de ces mutualismes tripartites. Je me suis tout d’abord intéressé à définir la relation qui lie de façon mutualiste le champignon aux deux autres partenaires et à établir les conflits d’intérêts émanant de ces différentes interactions pour comprendre quels facteurs permettaient de les réguler. Ainsi, la relation qui lie le champignon aux fourmis peut être qualifiée d’agriculture. Les fourmis protègent, nourrissent et disséminent le champignon et celui-ci, via ses propriétés structurales, permet l’élaboration de galeries servant de piège pour capturer des proies. Ce phénomène crée cependant un conflit d’allocation de la force ouvrière des fourmis et nuit directement aux bénéfices de la plante par une diminution de l’intensité des patrouilles sur ses feuilles, diminuant consécutivement sa protection et sa fitness. Néanmoins, le rôle du champignon dans les transferts de nutriments entre les fourmis et la plante, ainsi que certaines réponses évolutives de la plante permettent de réguler ces conflits, stabilisant les bénéfices nets de chaque partenaire dans ce mutualisme tripartite. Puis, je me suis concentré à comprendre comment certains facteurs évolutifs pouvaient moduler cette résultante écologique. La prise en compte du caractère multipartite d’un mutualisme change radicalement la vision de l’évolution des mutualismes jusqu’alors étudiée entre paires d’espèces. Alors que le corpus scientifique s’accorde à dire que la spécialisation entre espèce par coévolution renforce la stabilité et les bénéfices perçus par chaque partenaire, le contexte multipartite semble altérer ces prédictions. Au contraire la spécialisation multipartite peut être dans certains cas un moteur d’instabilité et de baisse des bénéfices. Enfin cette thèse permet de faire le lien entre deux concepts qui s’opposent : la coévolution diffuse et la coévolution par paire. Je montre ainsi que la coévolution peut intervenir sur plus de deux espèces à la fois, mais qu’elle peut quand même entrainer une spécialisation multispécifique. Finalement, au contraire des prédictions de la coévolution diffuse, cette thèse montre que plusieurs pressions de sélection contrastées émanant de différentes espèces envers un seul trait d’une troisième espèce peuvent promouvoir la spécialisation multi-spécifique.
Mutualisms, defined as interspecific interactions where each partner receives net benefices from their interactions, are central to the organization of earth biodiversity. Although globally beneficial for each partner, such interactions do not modify the inherent selfishness of species for their survival and reproduction, generating conflicts of interests between species. Thus understanding the ecological and the evolutionary processes maintaining positive outcomes in mutualisms is fundamental to understand how mutualisms shape earth biodiversity. However, scientific research on mutualisms has most of the time focused on interaction between pairs of species. Such knowledge is thus partial as mutualisms are embraced in a community context. My doctoral thesis takes place in this conceptual framework. I focused my research on the integration of a third fungal partner in protective interactions between ants and plants to evaluate its consequences on the ecological and evolutionary outcomes of such mutualisms, taken as multispecies interactions. I first focused my researches in defining the mutualistic interaction linking the fungal partner with its two other associates and in revealing any conflict of interests and their regulation that may emerge from such tripartite interactions. The interactions between the ants and the fungi can be qualified as a case of non-food fungiculture. The ants protect, provide food and disseminate the fungus, and the latter, thanks to its structural properties, allows the elaboration of galleries used that are then used as trap to capture preys. This phenomenon creates a conflict of interest in the allocation of the worker force, altering host plant benefits through a decrease of worker patrolling activity and consequently leaves protection and thus fitness. However, the role of the fungus in the nutrient transfers between ants and plants added to evolutionary responses from the plant allows regulating this conflict, stabilizing the net benefits towards the plant. Then I have concentrated my researches in understanding how evolutionary factors would modulate the ecological outcomes of such interactions. Taking into account the multispecific character of mutualisms changes radically the vision on the evolution of mutualisms when pair of species are considered. While it is widely accepted that specialization of mutualist species through coevolution reinforce the stability of the interaction and the net benefit of each partner, the multispecific context seems to deviate these predictions. Conversely, I show that specialization between three mutualistic partners can drive instability and decrease of benefits. Finally, the results of this thesis join the gap between two previously opposed concepts: diffuse coevolution and pairwise coevolution. I show that coevolution can happen between more than two species simultaneously and that it can drive mutlispecific specialization. Opposed to the diffuse coevolution, I show that contrasting selective pressures on a same trait from different partner can promote specialization of species.

http://www.theses.fr/2013AGUY0622/document

Résumé : La gestion du foncier martiniquais est un sujet complexe, sensible et à forte densité politique. La superficie de la Martinique est de 1 100km2, soit l’une des plus petites régions françaises, avec la plus forte densité de population, soumise à de multirisques (cyclonique, sismique, volcanique, technologique, inondation, pollution de l’air, des sols et de l’eau, tsunami…).La triangulation urbanistique composée d’une multiplicité d’acteurs, de textes et de réglementations et d’un millefeuille institutionnel (6 niveaux d’administration : commune, intercommunalité, département, région, Etat, Europe) annihile toute responsabilité et obère un développement endogène de ce micro territoire.Un juste équilibre entre développement économique, social, culturel et environnemental doit être trouvé en conciliant les objectifs économiques et sociaux liés au développement insulaire.La remarquable biodiversité de la Martinique constitue un atout touristique indéniable, menacé par de fortes pressions anthropiques foncières.Cette recherche démontre la pertinence de la réglementation applicable au foncier à la Martinique et l’influence du développement durable sur cette réglementation. La thèse propose des solutions pérennes à mettre en œuvre pour pallier les incohérences juridiques révélées par l’étude.
Management of land in Martinique is a complex, sensitive and highly political topic.The total area of the island is 1 100 km2 (square meters), e.g, one of the smallest French areas, with the highest concentration of population, submitted to multirisk desasters (cyclonic, seismic, volcanic, technological, flood, air, ground and water pollutions, tsunami…).The urban triangulation made up of a multiplicity of actors, texts and regulations, as well as a multi-level institutional cream cake (6 levels of administration: the city, the intercity, the Department, the Regional Council, the State and Europe), destroys all kind of responsibility and obstruct an endogenous development of this micro territory.A fair balance between economic, social, cultural and environmental development is necessary, by reconciling with the economic and social goals related to the insular development.The astonishing biodiversity of the island of Martinique represents an obvious tourist asset, threatened by strong land anthropic pressures.This research study highlights the relevance of the regulation applicable to the land in Martinique as well as the influence of sustainable development on this regulation. This thesis suggests perennial solutions to implement, in order to mitigate the legal inconsistencies pointed out by the study.

http://www.theses.fr/2011AGUY0503/document